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Herzogia 28 (1), 2015: 28 –37

Calciphilous species of the genus Scapania in the Tatra Mountains (Western Carpathians, Poland and Slovakia): distribution and threats Piotr Górski & Jiří Váňa

Abstract: Górski, P. & Váňa, J. 2015. Calciphilous species of the genus Scapania in the Tatra Mountains (Western Carpathians, Poland and Slovakia): distribution and threats. – Herzogia 28: 28 –37. This study presents the distribution, both horizontal and vertical, of eight species of the genus Scapania occurring throughout the Tatra Mountains, on the strongholds of limestone rocks (S. aequiloba, S. aspera, S. brevicaulis, S. calcicola, S. cuspiduligera, S. gymnostomophila, S. helvetica, and S. praetervisa). A list of localities of each species is presented on the basis of the data published between 1814 and 2014 as well as our own herbarium data. An original key for identification of earworts was prepared, and selected features are also illustrated. Zusammenfassung: Górski, P. & Váňa, J. 2015. Kalkliebende Arten der Gattung Scapania in der Tatra (Westliche Karpaten, Polen und Slowakei): Verbreitung und Gefährdung. – Herzogia 28: 28 –37. In der vorliegenden Studie wird die horizontale und vertikale Verbreitung von acht kalkliebenden Arten der Gattung Scapania in der Tatra (S. aequiloba, S. aspera, S. brevicaulis, S. calcicola, S. cuspiduligera, S. gymnostomophila, S. helvetica und S. praetervisa) dargestellt. Für alle Arten wird die Verbreitung auf der Basis von zwischen 1814 und 2014 publizierten Literaturdaten und von eigenen Herbardaten dokumentiert. Neben einem Bestimmungsschlüssel werden ausgewählte Merkmale illustriert. Key words: Scapaniaceae, Tatra Mts, Western Carpathians.

Introduction Within the genus Scapania (Dumort.) Dumort., 24 species have been detected throughout the entire area of the Tatra Mts. The first study on the flora of liverworts occurring in the Tatra Mts (Wahlenberg 1814) lists merely Scapania undulata (L.) Dumort. in the whole group of 30 species. Before the end of the 19th century, 11 species from the Tatra Mts had already been identified. This was possible due to the first monograph of liverworts of the area (Szyszyłowicz 1885), but also to shorter floristic notes (Rehman 1864, Limpricht 1877a, Krupa 1882, 1888). The localities of very rare earworts were also identified in that period among others. They include S. apiculata Spruce or S. compacta (Roth) Dumort. The number of their localities in the Tatra Mts does not exceed 10 at the moment. In the 20th century, the first monographic studies in the Scapania genus appeared. Those worth mentioning include the Polish monograph of the genus Scapania (Hausbrandt 1949) covering the whole area of Poland (including the Polish Tatras but also West Ukraine and Belarus). Single localities of mountain species from the Slovak Tatra Mountains can also be found in the paper. The study encompasses the first records of Scapania crassiretis Bryhn or S. glaucocephala (Taylor) Austin var. scapanioides (C.Massal.) Potemkin [= S. scapanioides (C.Massal.)

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Grolle] in Poland and the Tatra Mts. In the second part of the 20th century, the first specification of all known localities of earworts from the Slovak Tatras were already published. They also started to issue the first reviews of herbarium materials of the genus (Duda & Váňa 1968, 1969a, b, 1970a, b, 1971a, b, 1972). The group of the Tatra species reported the latest include Scapania aspera Bernet et M.Bernet (Duda & Váňa 1970a; the previous data are incorrect), S. praetervisa Meyl., S. gymnostomophila Kaal., S. parvifolia Warnst. (Szweykowski 1956a, b, 1960) and S. carinthiaca J. B.Jack ex Lindb. (Duda 1955a, b). Presently, S. undulata (L.) Dumort. is the most widespread in the whole area of the Tatra Mts (ca. 400 localities recorded between 1814 and 2013), and there are also S. aequiloba (Schwaegr.) Dumort. (ca 290), S. subalpina (Nees ex Lindenb.) Dumort. (ca 125), S. uliginosa (Sw. ex Lindenb.) Dumort. (ca 115) and S. umbrosa (Schrad.) Dumort. (ca 70) (Górski & Váňa 2014). One of the most influential factors of the distribution of plants in the Tatra Mts is their ground preference. Eight species within the genus Scapania grow on the strongholds of limestone rocks. They include Scapania aequiloba, S. aspera, S. brevicaulis Taylor, S. calcicola (Arnell et J.Perss.) Ingham, S. cuspiduligera (Nees) Müll.Frib., S. gymnostomophila, S. helvetica Gottsche and S. praetervisa. The objective of the following study is presenting the horizontal and vertical distribution of calciphilous species of the genus Scapania as well as determining their current threats in the massif. An original key for the determination of the species of this group is also presented.

Methods All known data regarding the occurrence of eight species of the genus Scapania that is S. aequiloba, S. aspera, S. brevicaulis, S. calcicola, S. cuspiduligera, S. gymnostomophila, S. helvetica and S. praetervisa have been arranged. This applies to the species occuring throughout the entire area of the Tatra Mts (both on the Polish and Slovak sides). The data include information published between 1814 and 2014 as well as the authors‘ own herbarium data collected in the Belianskie Tatra Mts and the limestone part of the Western Tatra Mts. Herbarium materials deposited in selected herbaria in Poland, Slovakia and the Czech Republic were also used (see list of localities). The degree of threat to the analyzed species throughout the Tatra Mts was specified with the use of categories and criteria of the IUCN (see IUCN 2001).

Area of research The Tatra Mountains are the highest mountain range in the whole Carpathians. They are a part of the Central-Eastern Carpathians. The length of the ridge of the Tatra Mts is close to 80 kms (56.5 kms in a straight line), while the maximum width is 18.5 kms (15 kms on average). The range is located on the border between Poland and Slovakia and covers an area of 785 km², including 610 km² (77.7 %) on the Slovak side and 175 km² (22.3 %) in Poland (RadwańskaParyska & Paryski 2004). The range is divided into: Belianskie Tatra Mts, High Tatra Mts and Western Tatra Mts. Sedimentary calciferous rocks cover mainly the northern part of the Tatras (Fig. 1). They, in turn, form the entire Belianskie Tatra Mts (= Belianskie Tatry Mts, Mt Havran, 2,154 m a.s.l.). The major limestone formations in the area of Poland in the Western Tatra Mts (= Tatry Zachodnie Mts) are formed by Mt Giewont (1,909 m a.s.l.), Kominiarski Wierch (1,829 m) and the northern parts of the mountain valleys (e.g., the Dolina Strążyska, Dolina Kościeliska, Dolina Chochołowska valleys). In the area of the Slovak Western Tatra Mts (= Západné

30 Herzogia 28 (1), 2015

Tatry Mts), there is Mt Osobitá (1,687 m), Mt Sivý vrch (1,806 m) and the surroundings of the Bobrovecká dolina valley and the Juráňova dolina valley. The most significant limestone formations along the Polish-Slovak border include the Czerwone Wierchy Massif (Červené vrchy; as far as Mt Krzesanica / Kresanica, 2,123 m), Mt Bobrowiec (Bobrovec, 1,663 m) as well as Mt Furkaska (Veľka Furkaska, 1,491 m).

Fig 1: The Tatra Mountains – area of investigation: 1 – Mt Gerlachovský štít (Gerlach; 2,655 m a.s.l.), 2 – Mt Rysy (2,499 m a.s.l.), 3 – a rough range of sedimentary calciferous rocks located in the northern part of the massif, 4 – state border.

Results Key to the species 1 Leaf cells with only 1(–2) large, 8 –15 µm wide, persistent oil-body in each cell. Plants small, to 1.5 cm long and 1–2 mm wide. Leaves keeled to base, unequally bilobed, both lobes entire, not decurrent, apex of lobes acute, apiculate to rounded; dorsal lobe small, 0.2– 0.4 the size of ventral lobe, patent to suberect, ventral lobe patent to squarrose. Keel 0.3 – 0.5 ventral lobe length. Marginal cells often smaller than median cells with thin or thickened walls. Cuticle smooth to obscurely papillosestriate. Gemmae common, dark brown to dark reddish brown, occasionally pale brown to yellow brown. ..................................................................................... S. gymnostomophila (sect. Calcicolae) 1* In each leaf cell more than one oil-body, persistent or mostly not persistent ............................. 2 2 Leaves not conspicuously decurrent ventrally or only decurrent to the level of insertion of the keel, decurrent strip only 1 cell wide. Cuticle smooth to papillose. Gemmae brown to reddish, at least becoming brownish (only young gemmae green) .............................................................. 3 2* Leaves distinctly decurrent ventrally below level of keel insertion, with the decurrent strip several cells wide. Cuticle smooth to coarsely papillose-striate. Gemmae mostly green, rarely yellowish to reddish brown, dark brown or red ............................................................................. 8 3 Leaves subequally bilobed, with dorsal lobe 0.7– 0.9 of the ventral in size. Leaves bordered by 2– 4 rows of cells with strongly thickened walls lacking trigones or marginal cells smaller than median cells and soon loosing oil-bodies ......................................................................................... 4 3* Leaves unequally bilobed, with dorsal lobe 0.3 – 0.7 of the ventral in size. Leaves not bordered by thick-walled cells (at least marginal cells only slightly smaller or slightly thick-walled, not loosing oil-bodies) ............................................................................................................................. 5 4 Leaves bordered by 2– 4 rows of cells with strongly thickened walls lacking trigones. Cuticle nearly smooth to weakly papillose-striate. Gemmae common, yellowish to reddish brown, dark brown or red. Plants 1.5 –2 cm long and 1–3 mm wide. Leaves often undulate, subequally bilobed, leaf bases sheathing. Leaf lobes similarly orientated and patent, erecto-patent to squarrose, obovo-

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id lingulate, apex mostly obtuse, rarely subapiculate to acute. Keel rounded, 0.5 – 0.7 ventral lobe length. Ventral leaf base formed of elongated thick-walled cells. Oil bodies large, 2–5 per cell, occupying up to 0.5 the cell lumen. .............................................. S. cuspiduligera (sect. Cuspiduligerae) 4* Marginal cells not with strongly thickened walls, only smaller than median cells and soon loosing oil-bodies. Cuticle very coarsely papillose. Gemmae green. . S. aequiloba (sect. Aequilobae) 5 Oil-bodies 2– 4(–5) per cell, large, persistent, up to 5 –15 × 5 –12 µm in diam., filling most of the cell lumen. Plants large (up to 5 cm) to smaller. ................................................................................ 6 5* Oil-bodies small, not persistent, 4 – 8 µm in diam., filling less than 0.5 of the cell lumen. Plants smaller, up to 1.5 cm long ................................................................................................................... 7 6 Plants large, 2–5 cm long. Dorsal lobe 0.8 –1.2 wider as long; ventral lobe broadly ovate to suborbicular, their width mostly 0.8 –1.25 their length; keel (at least on some leaves) strong­ ly semicircularly arched. Gemmae mostly 1-celled, green to brown. Leaves unequally bilobed, with dorsal lobe erecto-patent to suberect and ventral lobe patent to erecto-patent, lobes mostly not decurrent (ventral lobe sometimes narrowly decurrent to below level of keel base), apex of lobes pointed to subacute or rounded. Dorsal lobe 0.5 – 0.8 of the size of ventral lobe, sometimes dentate. Keel 0.3 – 0.5 ventral lobe length. Cuticle smooth to weakly verrucose. Mostly on humus-rich soil on limestone. .......................................................................................... S. brevicaulis (sect. Nemorosae) 6* Plants smaller, up to 2.5 cm long and 1–3 mm wide. Dorsal lobe 0.6 – 0.9 wider as long; ventral lobe narrow, their width 0.4 – 0.8 their length; keel straight to slightly arcuate. Gemmae mostly 2-celled, green, gradually becoming pale yellow-brown to brown. Leaves unequally bilobed with dorsal lobe patent and ventral lobe patent to squarrose, lobes not or shortly decurrent, apex of lobes rounded to acute or apiculate. Dorsal lobe 0.4 – 0.6 of the size of ventral lobe, sometimes distantly dentate. Keel 0.3 – 0.6 ventral lobe length. Marginal cells usually slightly thick-walled. Cuticle smooth to weakly papillose. On limestone or dolomite rocks. ............. S. calcicola (sect. Calcicolae) 7 Dorsal lobe dilated above base, obtuse to subapiculate, convex, ventral lobe always obtuse. Leaf margins entire. Perianth mouth subentire to only finely remotely denticulate by small 1–2 celled teeth. Plants 1–1.5 cm long and 1–2.5 mm wide, weakly to shortly decurrent. Keel 0.5 – 0.7 ventral lobe length. Marginal cells not or only slightly smaller than the median ones. Cuticle smooth to papillose. Gemmae not common, green to slightly brown. .................... S. helvetica (sect. Curtae) 7* Dorsal lobe acute to apiculate, erecto-patent, mostly flat, about 0.5 – 0.75 to nearly as long as the patent, weakly concave ventral lobe with rounded to apiculate apex. Leaf margins often denticulate. Perianth mouth laciniate with up to 8 cells long cilia. Plants 1–1.5 cm long and 1–2.5 mm wide. Keel 0.5 – 0.8 ventral lobe length. Marginal cells not or only slightly smaller than the median ones. Cuticle smooth to papillose. Gemmae not common, green to slightly brown, mostly reddish brown. ..................................................................................................... S. praetervisa (sect. Curtae) 8 Cuticle smooth to weakly or moderately papillose or verrucose .................................................. 9 8* Cuticle very coarsely papillose with hemispherical papillae. Gemmae 2– 4-celled, green to brown ............................................................................................................................................... 10 9 Leaves subequally bilobed; keel ± straight. Gemmae common, mostly 2-celled, yellowish to reddish brown, dark brown or red. On limestone or dolomite rocks. .......... S. cuspiduligera (sect. Cuspiduligerae) 9* Leaves unequally bilobed; keel strongly semicircularly arched. Gemmae mostly 1-celled, green. On humus-rich soil in limestone. ...................................................... S. brevicaulis (sect. Nemorosae) 10 Branching lateral. Leaf lobes subequal, standing stiffly away from each other, rounded or apiculate, similarly orientated and erecto-patent; dorsal lobe plane to slightly convex, not decurrent, recurved or squarrose (Fig. 2f), ventral lobe plane or convex to strongly decurved, only slightly decurrent at base. Leaf margin edentate or distally dentate (Fig. 2g). Marginal cells smaller than median cells (Fig. 2h) and soon loosing oil-bodies. Plants to 4 –5 cm long and 1–3.5 mm wide. Keel 0.3 – 0.5 ventral lobe length. Gemmae common, 20 –30 µm long. ..................... ............................................................................................................. S. aequiloba (sect. Aequilobae)

32 Herzogia 28 (1), 2015

10* Branching exclusively ventral. Leaf lobes unequal, lying loosely over each other, apiculate, patent to erecto-patent; dorsal lobe weakly to strongly convex, ½–¾ of the size of ventral lobe, shortly decurrent, ventral lobe plane to slightly convex (Fig. 2b), decurrent at base. Leaf margin subdentate, dentate to dentate-serrate (Fig. 2c). Marginal cells scarcely smaller than median cells (Fig. 2d) and containing oil-bodies. Plants to 6(–9) cm long and 1– 4.5 cm wide. Keel 0.2– 0.4 ventral lobe length. Gemmae not so common, 25 – 40 µm long. ................S. aspera (sect. Aequilobae)

A

B

E

F

C

G

D

H

Fig 2: Scapania aspera (A–D) and S. aequiloba (E–H): A, B, E, F – part of leafy shot, dorsal side (bars: A – 10 mm, B – 2,5 mm, E – 12 mm, F – 4 mm); C, G – leaf margin (ventral lobe); D, H – marginal and median cells (dorsal lobe; bars C–D – 50 mm). All from the Tatra Mts; phot. A, B, E, F – M. Romański, C, G, D, H – P. Górski.

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Distribution of the species Amongst the eight species of Scapania occuring on the strongholds of limestone rocks in the Tatra Mts, the most widespread are Scapania aequiloba (292 localities) and S. cuspiduligera (99). One of the most common species is also S. helvetica (139). Most of its localities are, however, found in the non-limestone part of the massif, mostly in the High Tatra Mts. The species that can be considered rare include S. calcicola (26) and S. brevicaulis (24). It is worth mentioning that S. calcicola occurs solely on limestone rocks (mainly in the Belianskie Tatras), while most of the localities of S. brevicaulis are located in the crystalline part of the Tatra Mts In the High Tatra Mts, S. gymnostomophila, so far considered very rare (Szweykowski 1958b), seems to have increased in numbers. Many localities of the species have been identified in the Belianskie Tatra Mts, including the peak of the highest mountain (Mt Havran, 2,154 m a.s.l., leg. P. Górski, 2012). Its further localities have been found on limestone rocks in the Western Tatra Mts and also on low rocks in the valley, which is lower than 1,000 m a.s.l. The rarest species in the group are Scapania praetervisa (three localities) and S. aspera (six localities). Various revised herbar data suggest the latter to have been mistaken for S. aequiloba. Following the criteria of IUCN, the species that are considered endangered in the area of the Tatra Mts include S. praetervisa (category: EN – endangered) and S. aspera (VU – vulnerable). The other species are not endangered (Tab. 1). Within the group, two liverworts (S. brevicaulis and S. calcicola) are uncommon and may be considered endangered in the future (category: NT – near threatened). Table 1: The number of localities, the range of height occurrence and threats of calciphilous species in the genus Scapania (Dumort.) Dumort. in the Tatra Mountains. Explanations: ↓ - species minimum, ↑ - species maximum. * IUCN (2001): EN – endangered, LC – least concern, NT – near threatened, VU – vulnerable; for B1a, B2a see criteria in IUCN (2001). Belian-T – Belianskie Tatra Mts; High-T_PL – High Tatra Mts, Polish part; High-T_SL – High Tatra Mts, Slovakian part; West-T_PL – Western Tatra Mts, Polish part; West-T_SL – Western Tatra Mts, Slovakian part.

Species / Area S. aequiloba

Belian-T

High-T_SL High-T_PL West-T_SL West-T_PL

15 104 ↓800 –2062 1000 –2300↑

19 910 –1560

18 ↓800 –1790

136 880 –1830

2 4 1325 –1500↑ ↓970 –1330

Threats*/Number of new localities LC/123

-

-

-

S. brevicaulis

6 1550 –2062

11 1700 –1950

1 2100↑

5 1500 –1700

1 ↓1480

NT/─

S. calcicola

13 ↓900 –2062↑

-

-

4 1491–1610

9 920 –1340

NT/8

S. cuspiduligera

10 4 64 4 ↓1030 –2154 1665 –2040 1393 –2307↑ 1500 –1645

18 1040 –1960

LC/26

S. gymnostomophila

16 1480 –2154

7 ↓965 –1760

LC/14

S. helvetica

27 70 13 1240 –2100 1560 –2633↑ 1000 –2200

13 ↓900 –2187

16 980 –1960

LC/28

1 1700↑

1 1685

EN (B1a)/2

S. aspera

S. praetervisa

1 ↓1480

3 3 6 1665 –1735 1700 –2307↑ 1555 –1687

-

-

VU (B2a)/5

34 Herzogia 28 (1), 2015

Fig. 3: Altitudinal distribution of Scapania (Dumort.) Dumort. species in the Tatra Mts at intervals of 100 metres.

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Two distinct subgroups can be distingiushed in relation to the horizontal and vertical distribution of the genus Scapania discussed in this study. These are alpine species (most of the localities are found in the sub-alpine and alpine zones). They include: Scapania brevicaulis, S. cuspiduligera, S. gymnostomophila, S. helvetica and S. praetervisa. The other group, so called mountainwide species, consists of plants distributed quite evenly on all the mountain belts. They include S. aequiloba, S. calcicola and probably S. aspera (Fig. 3). A list with all localities of the investigated species in the Tatra Mts is given in the online supplement (Supplementary Table S1).

Acknowledgements Photos of Scapania aequiloba and S. aspera for this paper were done by Maciej Romański (Wigry National Park) whose help is gratefully acknowledged.

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Manuscript accepted: 07 November 2014. Communicated by: Harald Kürschner

Addresses of the authors Piotr Górski, Department of Botany, Poznań University of Life Sciences, Wojska Polskiego 71c, 60-625 Poznań, Poland. E-mail: [email protected] Jiří Váňa, Department of Botany, Charles University, Benátská 2, 128 01 Praha 2, Czech Republic. E-mail: [email protected]

Supplementary documents online: Supplementary Table S1: List of localities. www.bioone.org/toc/heia/28/1

GÓRSKI, P. & VÁŇA, J. 2015. Calciphilous species of the genus Scapania in the Tatra Mountains (Western Carpathians, Poland and Slovakia): distribution and threats. – Herzogia 28: 28–37. Supplementary Table S1: List of localities. Scapania aequiloba (Schwaegr.) Dumort. Literature data WESTERN TATRA MTS., Poland: Dolina Wyżnia Chochołowska valley, on shady rocks near Potok Siwa Woda stream, alt. 1320 m, leg. J. Szweykowski, 28.07.1957 (Szweykowski 1960); Dolina Chochołowska valley, on wayside limestone rocks by the valley estuary, alt. 935 m, leg. J. Szweykowski, 19.08.1956 (SZWEYKOWSKI 1960); Dolina Chochołowska valley, on a wayside limestone rock opposite Mt Zawiesista Skała, alt. 1025 m, leg. J. Szweykowski, 19.08.1956 (SZWEYKOWSKI 1960); Dolina Chochołowska valley, Polana Huciska glade (BALCERKIEWICZ & BRZEG 1978); Mt Kominiarski Wierch, humus between limestone rocks under the peak, alt. 1800−1825 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, cracks of limestone rocks on the western slope, alt. 1800 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, humus between limestone rocks on N slope above Dolina Smytnia valley, alt. 1750 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, humus between limestone rocks on the southern slope above Zdrapiska, alt. 1745 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1960); Dolina Kościeliska valley, on wayside rocks near Hala Pisana glade, alt. 1040 m, leg. J. Szweykowski, 22.08.1956 (SZWEYKOWSKI 1960); Dolina Kościeliska valley, damp limestone rocks in Wąwóz Kraków gully, alt. 1100 m, leg. J. Szweykowski, 29.06.1948 (SZWEYKOWSKI 1960); Dolina Kościeliska valley, Wąwóz Kraków gully, alt. 1035 m (SZYSZYŁOWICZ 1885); Dolina Kościeliska valley, Wąwóz Kraków gully, leg. F. Lilienfeldówna, 8.09.1912 (LILIENFELDÓWNA 1914a; KRAM 44167, WA); Dolina Kościeliska valley, alt. 939−1389 m (KRUPA 1882, SZWEYKOWSKI 1958a); on the bank of Kirowa Woda stream, leg. F. Lilienfeldówna, T. Wilczyński, 17.08.1912 (LILIENFELDÓWNA 1914a; KRAM 44164, WA); Dolina Miętusia valley, on shady limestone rocks in the spruce forest Wantule, alt. 1170 m, leg. J. Szweykowski, 19.07.1948 (SZWEYKOWSKI 1960); Dolina Małej Łąki valley, alt. 1209 m (KRUPA 1882, SZWEYKOWSKI 1958a); Dolina Małej Łąki valley, humus between limestone rocks on a rocky treshold, alt. 1745 m, leg. J. Szweykowski, 11.09.1955 (SZWEYKOWSKI 1960); Dolina Strążyska valley, alt. 893-1040 m (KRUPA 1882, SZWEYKOWSKI 1958a); Dolina Strążyska valley, alt. 1040 m (SZYSZYŁOWICZ 1885); Dolina Strążyska valley, above Siklawica waterfall, leg. F. Lilienfeldówna, T. Wilczyński, 1.08.1912 (LILIENFELDÓWNA 1914a; KRAM 44166, WA); Dolina Strążyska valley, on damp limestone rocks, alt. 1000 m, leg. J. Szweykowski, 19.07.1948 (SZWEYKOWSKI 1960); Dolina Białego valley, alt. 941-1079 m (SZYSZYŁOWICZ 1885); Dolina ku Dziurze valley, alt. 894-1026 m (SZYSZYŁOWICZ 1885); Dolina Kondratowa valley, humus between limestone rocks into Piekło, alt. 1550 m, leg. J. Szweykowski, 9.09.1955 (SZWEYKOWSKI 1960); Mt Giewont, in cracks of limestone rocks on the southern slope, alt. 1830 m, leg. J. Szweykowski, 9.09.1955 (SZWEYKOWSKI 1960); Mt Giewont, Przełęcz Szczerba pass, humus between limestone rocks, alt. 1825 m, leg. J. Szweykowski, 9.09.1955 (SZWEYKOWSKI 1960); Mt Mały Giewont, alt. 1728 m, leg. J. Żebrowska (HAUSBRANDT 1949; WA); Wrótka under Giewont, alt. 1079-1428 m (SZYSZYŁOWICZ 1885); Dolina Jaworzynki valley, leg. J. Żebrowska (HAUSBRANDT 1949; WA); Mt Nosal, alt. 980-1215 m (SZYSZYŁOWICZ 1885); Mt Nosal, alt. 1206 m, leg. J. Żebrowska (HAUSBRANDT 1949; WA); Mt Kopa Magury, near Jaskinia w Magurze cave, alt. 1460 m (SZYSZYŁOWICZ 1885); Jaskinia w Magurze cave, alt. 1400 m (SCHIFFNER 1908); between Mt Kopa Magury and Mt Gładkie Jaworzyńskie, alt. 1460−1600 m (SZYSZYŁOWICZ 1885); Mt Czerwone Wierchy (KRUPA 1882); Mt Smreczyński Wierch, leg. J. Krupa (HAUSBRANDT 1949); WESTERN TATRA MTS, Slovakia: Račkova dolina valley, alt. 800 m (ŠMARDA 1937, DUDA 1955a, DUDA & VÁŇA 1970a); below Mt Tomanová, alt. 1440 m, leg. A. Zlatník, 1964 (DUDA & VÁŇA 1970a); Tomanová dolina valley, Žleb spod Diery gully, alt. 1535 m (UNAR et al. 1984−1985); Suchá dolina valley, below Mt Osobitá, alt. 1500 m (SZYSZYŁOWICZ 1885, HAUSBRANDT 1949); Mt Suchý Hrádok, alt. 1200 m (ŠMARDA 1937, DUDA 1955a); Diera, alt. 1500 m (ŠMARDA 1961); Rozpadlá dolinka valley (Rozpadliny), alt. 1690 m (ŠMARDA 1961); Mt Sivý vrch, alt. 1600 m (PECIAR 1981); HIGH TATRA MTS, Poland: Dolina Roztoki valley, leg. L. Goetzen-Hausbrandt (HAUSBRANDT 1949; WA); Mt Wołoszyn, leg. J. Krupa (HAUSBRANDT 1949; WA); Łysa Skałka near Łysa Polana, alt. 1025 m, leg. H. Klama (KLAMA 2008); HIGH TATRA MTS, Slovakia: slopes down the peak Jahňací štít, above Kolový potok stream, alt. 1520 m, leg. A. Zlatník, 1964 (DUDA & VÁŇA 1970a); Tatranská Javorina, alt. 993 m (LIMPRICHT 1877a, b, SZYSZYŁOWICZ 1885); Mengusovská dolina valley, Červený žľab gully, alt. 1610 m (ŠMARDA 1960b, 1961); between Hincova plesa lakes and Kôprovské sedlo pass, alt. 2040 m (ŠMARDA 1961); Zlomisková štrbina, alt. 2270-2300 m (ŠMARDA 1961); Ťažká dolina valley, alt. 1650 m, leg. L. Goetzen-Hausbrandt (HAUSBRANDT 1949); Bielovodská dolina valley (SZYSZYŁOWICZ 1885, HAUSBRANDT 1949; WA); Bielovodská dolina valley, Mt Holica (ŠMARDA 1961); Javorová dolina valley, alt. 1000 m (ŠMARDA 1961); Zlomisková dolina valley, Zlomiskova roveň, alt. 1810 m (ŠOLTÉS 2004); BELIANSKIE TATRA MTS: Mt Javorinka, leg. G. Limpricht, 11.07.1874 (DUDA & VÁŇA 1970a); Mt Javorinka, alt. 1000−1200 m, leg. A. Boros, 21.08.1962 (DUDA & VÁŇA 1970a); above Podspády, Javorinka stream (LIMPRICHT 1877a, 1877b, SZYSZYŁOWICZ 1885, HAUSBRANDT 1949); Mt Muráň, alt. 1200, leg. A. Boros, 17.08.1963 (DUDA & VÁŇA 1970a); Mt Nový vrch, alt. 1200 m, leg. A. Boros, 17.08.1963 (DUDA & VÁŇA 1970a); Mt Ždiarska vidla, humus between limestone rocks on the northern slope under the peak, alt. 2000 m, leg. J. Szweykowski, 19.07.1956 (SZWEYKOWSKI 1960); Mt Ždiarska vidla, on damp limestone rocks on the western slope above the path, alt. 1950 m, leg. J. Szweykowski, 19.08.1956 (SZWEYKOWSKI 1960); Mt Havran, alt. 1300 m, leg. L. Vajda, 12.07.1962 (DUDA & VÁŇA 1970a); Mt Havran, leg. A. Boros, 2.09.1955, 21.08.1962 (DUDA & VÁŇA 1970a); Mt Havran, alt. 1500−1600 m, leg. A. Boros, 31.08.1962 (DUDA & VÁŇA 1970a); Havrania dolina valley, alt. 1700 m, 1750 m (SUZA 1937, ŠMARDA 1938); between Mt Havran and Mt Ždiarska vidla, alt. 1325 m, leg. V. Krajina, 19.08.1925, det. J. Duda (DUDA & VÁŇA 1970a); Podspády, below Mt Havran, leg. G. Limpricht, 12.07.1874 (DUDA & VÁŇA 1970a); Rígľanská dolina valley, alt. 1000−1300 m, leg. A. Boros, 25.08.1963 (DUDA & VÁŇA 1970a); Monkova dolina valley, alt. 900−950, leg. A. Boros, 25.08.1963 (DUDA & VÁŇA

 

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1970a); Predné Meďodoly valley, leg. Z. Pilous, 21.06.1938 (DUDA 1958; DUDA & VÁŇA 1970a); Široké pole, leg. F. Grüll, 1950 (DUDA 1962; DUDA & VÁŇA 1970a); Široké pole, alt. 1700 m (ŠMARDA 1938); Mt Hlúpy vrch, alt. 2062 m, leg. J. Šmarda, 18.07.1937 (ŠMARDA 1938, 1961; DUDA & VÁŇA 1970a); Mt Hlúpy vrch, alt. 2050, leg. J. Váňa, 5.09.1967 (DUDA & VÁŇA 1970a); Mt Hlúpy vrch, alt. 2062 m (BOROS et al. 1960); between Mt Hlúpy vrch and Mt Zadné Jatky, alt. 1900 m, leg. J. Váňa, 5.09.1967 (DUDA & VÁŇA 1970a); Mt Zadné Jatky, on the humus of the snow-beds on the northern slope, alt. 1800 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Zadné Jatky, alt. 2000 m, leg. A. Boros, 15.07.1958 (DUDA & VÁŇA 1970a); Mt Bujači vrch, N slope, alt. 1300 m, slope E, alt. 1500−1600 m (SCHIFFNER 1911); Mt Bujači vrch, alt. 1800 m (ŠMARDA 1956); Mt Bujači vrch, alt. 1950 m, leg. J. Šmarda, 19.07.1937 (DUDA & VÁŇA 1970a); between Mt Bujači vrch and Mt Kopský hrebeň, alt. 1600 m, leg. J. Podpěra, 21.07.1922 (DUDA & VÁŇA 1970a); Babi potok stream, alt. 1000 m, leg. I. Györffy, 4.09.1911 (DUDA & VÁŇA 1970a); Skalné vráta pass, on a limestone rock, on the southern slope, alt. 1610 m, 1620 m, leg. J. Szweykowski, 18.07.1956, 21.07.1956 (SZWEYKOWSKI 1960); Skalné vráta pass, alt. 1300-1600 m, leg. A. Boros, 7.07.1933 (DUDA & VÁŇA 1970a); Skalné vráta pass, alt. 1600 m, leg. J. Podpěra, 29.07.1922 (DUDA & VÁŇA 1970a); Skalné vráta pass, leg. J. Šmarda, 8.1947 (ŠMARDA et al. 1962, DUDA & VÁŇA 1970a); Skalné vráta pass, above chalet Plesnivec, alt. 1600 m (DUDA 1955a); Dolina Siedmych prameňov valley, calcareous rocks near Chata Plesnivec, leg. L. Goetzen-Hausbrandt (HAUSBRANDT 1949; WA); Alabastrová jaskyňa cave, alt. 1300−1400 m, leg. A. Boros, 24.08.1938 (BOROS 1942 sub S. aspera, BOROS 1951, DUDA & VÁŇA 1970a); Mt Kozi chrbát, on the slope of the path between Sedlo Pod Schody pass and Mt Faixová, alt. 1500 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Kozi chrbát, on the slope of the path between Skalné vráta pass and Sedlo Pod Schody pass, alt. 1600 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); Mt Kozi chrbát (ŠMARDA 1958); Mt Kozi chrbát, Mt Limbová veža, Mt Jelenia skála (ŠMARDA et al. 1962); Mt Kozi chrbát, Mt Velká Jelenia skála (ŠMARDA 1961); Sedlo Pod Schody pass, on the damp and shady limestone rocks by the path under the pass, alt. 1500 m, leg. J. Szweykowski, 18.07.1956 (SZWEYKOWSKI 1960); Faixová lúka, leg. A. Vĕzda, 13.07.1949 (DUDA & VÁŇA 1970a); Červená hlina, alt. 1100 m, leg. I. Györffy, 29.06.1910 (SCHIFFNER 1911); Červená hlina, leg. A. Boros sub S. aspera, rev. J. Duda (BOROS 1951; DUDA & VÁŇA 1970a); Ždiar: Spod Šiky, alt. 1100 m (ŠMARDA 1961); Mt Bednárský, alt. 1000 m (ŠMARDA 1938); Malý Košiar, alt. 1900 (ŠMARDA 1961); Plesnivec, leg. L. Goetzen-Hausbrandt (HAUSBRANDT 1949); Modřinová dolina valley, alt. 1100 m, leg. J. Podpěra, 29.07.1922 (DUDA & VÁŇA 1970a); Čierny potok stream, in the stream valley, leg. F. Hazslinszky (DUDA & VÁŇA 1970a); Čierny potok stream, in the stream valley, alt. 900−1300 m, leg. A. Boros, 7.07.1933 (DUDA & VÁŇA 1970a); Čierny potok stream, in the stream valley, alt. 1270 m, leg. I. Györffy, 4.07.1911 (DUDA & VÁŇA 1970a); Dolina siedmich prameňov valley (JEŽEK & VONDRÁČEK 1962); Tatranská Kotlina, alt. 800 m, leg. J. Podpěra, 26.07.1922 (DUDA & VÁŇA 1970a); Belianskie Tatra (precise location not given), leg. T. Wiśniewski (HAUSBRANDT 1949); TATRA MTS, precise location not given (REHMAN 1864). New localities WESTERN TATRA MTS, Poland: Dolina Chochołowska valley, at the mouth of the Koziniecki Żleb gully, alt. 880 m, leg. P. Górski, 18.08.2005 (POZNB 368, KRAM-B); Dolina Chochołowska valley, near Wywierzysko Chochołowskie well-head, on limestone rocks, alt. 990 m, leg. P. Górski, 29.09.2003 (POZNB 377); Dolina Chochołowska valley, near Wyżnia Chochołowska Brama, outcrops of limestone rocks, alt. 1030 m, leg. P. Górski, 8.08.2004 (POZNB 443); Dolina Chochołowska valley, Sucha Dolina Wielka valley, a boulder on the edge of the road, alt. 980 m, leg. J. Szweykowski, 24.09.1992 (POZW 33337); Jasiorów Żleb gully, on the rock, alt. 1475 m, leg. J. Szweykowski, 27.08.1987 (POZW 252); Jasiorów Żleb gully (in the lower part), above Polana Chochołowska glade, on a limestone boulder, alt. 1230 m, leg. P. Górski, 8.08.2004 (POZNB 376); Dolina Dudowa valley, Między Ściany, on a limestone ledge, Caricetum firmae, alt. 1050 m, leg. P. Górski, 29.09.2003 (POZNB 444); Wielkie Koryciska valley, in the western part, on limestone walls near the stream, alt. 980 m, leg. P. Górski, 29.09.2003 (POZNB 366, KRAM-B ); Wielkie Koryciska valley, a clay and limestone slope near the stream, alt. 955 m, leg. P. Górski, 18.09.2005 (POZNB 383); Wielkie Koryciska valley, on eroded soil near the stream, alt. 1040 m, leg. J. Szweykowski, 23.08.1987 (POZW 246); Siwiańskie Turnie, on the humus of limestone rocks in the forest, alt. 990 m, leg. J. Szweykowski, 21.09.1989 (POZW 28115); Mącicki Żleb gully, above Siwa Polana glade, a damp stone in the forest near the stream, alt. 980 m, leg. J. Szweykowski, E. Chudzińska, 20.08.1993 (POZW 34515); rocks on the NE slope of Mt Wołowiec, between Skrzynia and Skrajniak, in a rock crack, alt. 1640 m, leg. J. Szweykowski, 21.08.1987 (POZW 64); Dolina Jarząbcza valley, rocks on the W slope of Mt Kończysty Wierch, alt. 1680 m, leg. J. Szweykowski, 18.08.1987 (POZW 7); Dolina Kościeliska valley, limestone rocks by the trail southwards from Polana Pisana glade (on the way to the hostel on Hala Ornak), alt. 1035 m, leg. P. Górski, 29.08.2008 (POZNB 372, KRAM-B); Dolina Kościeliska valley, by the trail on Polana na Stołach glade, 49°15.269'N, 19°51.703'E, alt. 1040 m, leg. P. Górski, 20.11.2011 (POZNB 757); Polana na Stołach glade, limestone rocks amongst meadows, 49°14.977'N, 19°51.362'E, alt. 1340 m, leg. P. Górski, 20.11.2011 (POZNB 756, 762, 800); Dolina Kościeliska valley, Wąwóz Kraków gully (in the part closed to visitors), alt. 1045 m, 1060 m, leg. P. Górski, 21.09.2005 (POZNB 373, 378, 379); Dolina Kościeliska valley, rocks at the bottom of Mt Kominiarski Wierch, Tylkowiańskie Spady, alt. 1505 m, leg. J. Szweykowski, 6.09.1990 (POZW 30634, 30066, 30630); Dolina Kościeliska, Pisaniarski Żleb gully, on a limestone boulder on the stream, alt. 1075 m, leg. J. Szweykowski, 3.09.1993 (POZW 34705); Dolina Kościeliska valley, at the mouth of the Pisaniarski Żleb gully, leg. K. Buczkowska, A. Bączkiewicz, 4.09.2006 (POZW 41098); Dolina Kościeliska valley, a slope southwards from the pass between Stoły and Mt Kominiarski Wierch, the edge of the path, alt. 1460 m, leg. J. Szweykowski, 6.09.1990 (POZW 30635); Mt Kominiarski Wierch, E part of Tylkowiańskie Spady, on the rocks, alt. 1575 m, leg. J. Szweykowski, A. Bączkiewicz, 1.09.1995 (POZW 37052); on a limestone rock by the trail Droga Nad Reglami, alt. 1150 m, leg. J. Szweykowski, 1.09.1989 (POZW 27523); near route from Dolina Kościeliska valley to Iwaniacka Przełęcz pass, Polysticho-Piceetum, alt. 1275 m, leg. P. Górski, 2.07.2004 (POZNB 1310); Dolina Miętusia valley, Krowi Żleb gully, under Upłaziński Wierszyk, humus on the slope, 4.09.2006, leg. K. Buczkowska, A. Bączkiewicz (POZW 41088, 41099, 41100); Dolina Miętusia valley, on the slope Adamica, on the stream, leg. J. Szweykowski, 14.09.1990 (POZW 30611); Dolina Miętusia valley, Krowi Żleb gully, under Upłaziński Wierszyk, humus on the slope, alt. 1040 m, leg. J. Szweykowski, 14.09.1990 (POZW 30532); Dolina Głębowiec valley, under Mt Bobrowiec, dolomite rocks, a stream temporarily dried up, on a rock, alt. 1060 m, leg. J. Szweykowski, 13.08.1988 (POZW 124/88); Dolina Lejowa valley,

 

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Między Ściany, on a rock, alt. 925 m, leg. J. Szweykowski, 24.09.1989 (POZW 27905); Dolina Potoku Spadowiec valley, on a stone in the streambed, alt. 1130 m, leg. J. Szweykowski, 7.09.1987 (POZW 310); Suchy Żleb Kalacki gully, alt. 1360 m, leg. J. Szweykowski, 3.08.1985 (POZW 23604); Dolina Kryta valley, limestone rocks on the stream, alt. 1025 m, leg. J. Szweykowski, 5.08.1989 (POZW 27337); Kuźnice, by the splitting of the trails into Boczań and Nosal, alt. 1085 m, leg. P. Górski, 16.09.2004 (POZNB 358, KRAM); Dolina Jaworzynka valley, Wielkie Siodło, limestone outcrops by the trail, alt. 1400 m, leg. P. Górski, 16.09.2004 (POZNB 283); Dolina Jaworzynka valley, Siodłowa Perć, below Przełęcz między Kopami pass, limestone rocks by the trail, alt. 1485 m, 49°15.278'N, E 020°00.016'E, leg. P. Górski, 30.08.2011 (POZNB 600); Dolina Jaworzynka valley, Wyżnia Rówień Jaworzynka, limestone rocks, Carici-Festucetum tatrae, alt. 1100 m, leg. P. Górski, 16.09.2004 (POZNB 814); Dolina Jaworzynka valley, above Polana Jaworzynka glade, a limestone boulder in the forest, near the trail, alt. 1175 m, leg. P. Górski, 16.09.2004 (POZNB 829); Dolina Bystrej valley, below Polana Kalatówki glade, by the trail to Mt Giewont, on a limestone boulder in a spruce forest Polysticho-Piceetum, alt. 1200 m, leg. P. Górski, 11.09.2004 (POZNB 442, KRAM); between Chłabówka and Cyrhla, calcareous rocks in the forest, near Oswald Balzer road, 49°16.998'N, 20°00.639'E, alt. 965 m, leg. P. Górski, 16.11.2012 (POZNB 1366); Żleb pod Czerwienicą gully, alt. 1510 m, leg. E. Chudzińska, J. Szweykowski, K. Buczkowska, 15.08.1998 (POZW 38455); Żleb pod Czerwienicą gully, alt. 1360 m, leg. J. Szweykowski, 15.07.1999 (POZW 38755); Dolina Olczyska valley, a slope under Pióro, on limestone rocks, alt. 950 m, leg. J. Szweykowski, 22.08.1993 (POZW 34479); Żleb Żeleźniak gully, a slope on the stream, alt. 1025 m, 7.09.1993, leg. J. Szweykowski, A. Bączkiewicz (POZW 34715); Mt Kończysta Turnia, alt. 1220 m, leg. J. Szweykowski, E. Chudzińska, 17.08.1993 (POZW 34480); Dolina Jaroniec valley (S from Polana Biały Potok glade), on a limestone rock by the road, alt. 1000 m, 16.08.1994, leg. E. Chudzińska, J. Szweykowski (POZW 36311); Żleb Wielki Ciemniak gully, alt. 1050 m, 1125 m, leg. J. Szweykowski, 22.08.1987 (POZW 226, 224a); Mt Wielka Świstówka, on limestone rocks, alt. 1370 m, leg. J. Szweykowski, 1.09.1987 (POZW 340); Potok Chłabówka stream, westwards of Zakopane, alt. 1000 m, on damp rocks of the closed quarry on the stream, leg. J. Szweykowski, 25.08.1985 (POZW 23770); in the middle part of the Kalackie Koryto (Suchy Żleb) gully, on a limestone boulder, in cracks, 49°15.357'N, 19°57.610'E, alt. 1335 m, leg. P. Górski, 24.11.2011 (POZNB 752, 779); in the SW part of Polana Kalatówki glade, high rock limestone walls in a spruce forest, at the mouth of Kalackie Koryto gully, 49°15.419'N, 19°57.754'E, alt. 1265 m, leg. P. Górski, 24.11.2011 (POZNB 791); Dolina za Bramką gully, at the mouth of the gully, limestone rocks in a beech forest, 49°16.661'N, 19°55.206'E, alt. 915 m, leg. P. Górski, 21.11.2011 (POZNB 769); Dolina za Bramką gully, high, vertical rock walls, 49°16.584'N, 19°55.159'E, alt. 920 m, leg. P. Górski, 21.11.2011 (POZNB 755); Dolina za Bramką valley, high, vertical rock walls, 49°16.566'N, 19°55.090'E, alt. 930 m, leg. P. Górski, 21.11.2011 (POZNB 795); Dolina za Bramką valley, shady, high rock walls in a spruce forest, above (westwards) the trail, down the main valley, 49°16.494'N, 19°54.971'E, alt. 990 m, leg. P. Górski, 21.11.2011 (POZNB 754); Dolina ku Dziurze valley, near Jaskinia Dziura cave, in the final part of the valley open to visitors, 49°16.318'N, 19°56.549'E, alt. 1005 m, leg. P. Górski, 21.11.2011 (POZNB 797); Dolina Małej Łąki valley, near the mouth of the valley, 49°16.543'N, 19°54.241'E, alt. 910 m, leg. P. Górski, 22.11.2011 (POZNB 765); Dolina Małej Łąki valley, just below the splitting of the trail into Przysłop Miętusi and Rówienki, 49°15.998'N, 19°54.056'E, alt. 1040 m, leg. P. Górski, 22.11.2011 (POZNB 778); Dolina Strążyska valley, a limestone rubble on the slope by the trail, at the mouth of the valley, 49°16.375'N, 19°55.978'E, alt. 935 m, leg. P. Górski, 21.11.2011 (POZNB 793); Dolina Strążyska valley, limestone rocks near the mouth of the valley, 49°16.255'N, 19°55.937'E, alt. 990 m, leg. P. Górski, 21.11.2011 (POZNB 776); Dolina Strążyska valley, a gully below (northwards) of Skała Jelinka, 49°16.162'N, 19°55.899'E, alt. 995 m, leg. P. Górski, 21.11.2011 (POZNB 794); Dolina Strążyska valley (precise location not given), ? (WA); Staników Żleb gully, at the mouth of the valley, shady, rock walls in a spruce forest, near stream, 49°16.168'N, 19°52.805'E, alt. 1075 m, leg. P. Górski, 22.11.2011 (POZNB 753, 792); Dolina Lejowa valley, a slope by the trail, above Polana Huty Lejowe glade, 49°15.481'N, 19°51.185'E, alt. 1050 m, leg. P. Górski, 20.11.2011 (POZNB 801); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), a small boulder in the lower part covered with humus, 49°15.157'N, 19°50.989'E, alt. 1145 m, leg. P. Górski, 20.11.2011 (POZNB 744, 761); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), in the lower part of the wetland, springs with Cratoneuron, 49°15.107'N, 19°50.972'E, alt. 1180 m, leg. P. Górski, 20.11.2011 (POZNB 760, 772); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), shady and vertical limestone walls of the gully, 49°15.103'N, 19°50.978'E, alt. 1180 m, leg. P. Górski, 20.11.2011 (POZNB 766, 773); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), in the middle part, on a stone near stream, 49°15.065'N, 19°50.960'E, alt. 1210 m, leg. P. Górski, 20.11.2011 (POZNB 767); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), a limestone wall in the middle part of the gully, 49°15.053'N, 19°50.945'E, alt. 1215 m, leg. P. Górski, 20.11.2011 (POZNB 764); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), in the middle part, 49°15.027'N, 19°50.934'E, alt. 1225 m, leg. P. Górski, 20.11.2011 (POZNB 768); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), two limestone boulders in the upper part filling the inside of the wide gully, on the humus and bare rock, 49°14.921'N, 19°50.860'E, alt. 1290 m, leg. P. Górski, 20.11.2011 (POZNB 759, 763); Kobylarzowy Żleb pod Małołączniakiem gully, in the lower part, rock walls, 49°14.624'N, 19°54.421'E, 49°14.602'N, 19°54.428'E, alt. 1505 m, 1540 m, leg. P. Górski, 22.11.2011 (POZNB 790, 789); Kobylarzowy Żleb under Małołączniak gully, in the upper part, rock walls, Caricetum firmae, on the humus or a bare rock wall, 49°14.632'N, 19°54.649'E, alt. 1700 m, leg. P. Górski, 22.11.2011 (POZNB 780); Kobylarzowy Żleb pod Małołączniakiem gully, in the upper part, limestone rocks, 49°14.668'N, 19°54.757'E, alt. 1760 m, leg. P. Górski, 22.11.2011 (POZNB 788); Dolina Gąsienicowa valley, above Przełęcz między Kopami pass, a limestone slope by the trail, alt. 1510 m, leg. P. Górski, 19.08.2005 (POZNB 816); Mały Żlebek valley, leg. F. Krawiec, 18.08.1928 (WA); Czerwona Przełęcz pass, near Mt Sarnia Skała, alt. 1301 m, leg. Rejment-Grochowska, 14.08.1957 (WA); WESTERN TATRA MTS, Slovakia: Bobrovecká dolina valley, alt. 900–1000 m, leg. J. Váňa, 30.05.1990 (herb. Váňa); Tichá dolina valley, alt. 950 m, leg. J. Váňa, 27.05.1990 (herb. Váňa); Juráňova dolina valley, Tiesňavy, alt. 1100 m, leg. J. Váňa, 25.05.1984 (herb. Váňa); Červené vrchy Massive, Javorový žlab gully, alt. 1700 m, leg. J. Váňa, 18.08.1995 (herb. Váňa); Mt Sivý vrch, slopes down the peak towards north, alt. 1790 m, leg. P. Górski, 19.07.2008 (POZNB 367); Jalovecká dolina valley, Mt Mních, alt. 1416 m, leg. R. Šoltés sub S. aspera, 30.08.1993, rev. P. Górski (Herbarium Múzeum Tatranského národnego parku, 1/7864); Juráňova dolina valley, Tiesňavy, calcareous outcrops in the lower part of the valley, 49°16.838'N, 19°46.293’E, alt. 860 m, leg. P. Górski, 30.09.2013 (POZNB 1664); Juráňova dolina valley, Válovec, calcareous outcrops in the forest, 49°15.344'N, 19°46.812'E, alt. 1270 m, leg. P. Górski, 30.09.2013 (POZNB 1677); HIGH

 

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TATRA MTS, Poland: Dolina Filipka valley, Potok Filipczański stream, on eroded soil, alt. 1100 m, leg. J. Szweykowski, 21.09.1988 (POZW 613/88); Dolina Filipka valley, Potok Filipczański stream, on a stone by a river bank, alt. 950 m, leg. J. Szweykowski, 21.09.1988 (POZW 611/88); Dolina Filipka valley, Potok Filipczański stream, on a rock by the road along the stream, alt. 1075 m, leg. J. Szweykowski, 21.09.1988 (POZW 629/88); Dolina Filipka valley, Żleb za Skałką gully, alt. 1075 m, leg. J. Szweykowski, 18.09.1992 (POZW 33305); Dolina Filipka valley, on Potok Filipczański stream, alt. 920 m, leg. J. Szweykowski, 22.09.1988 (POZW 751/88); Dolina Filipka valley, on a thin layer of humus, on a stone on the bank of Potok Filipczański stream, alt. 950 m, leg. J. Szweykowski, 22.09.1988 (POZW 757/88); Dolinka Skalnite valley, alt. 930 m, leg. J. Szweykowski, 16.09.1992 (POZW 33192); Koziarski Żleb gully, on a limestone rock on the stream, alt. 970 m, leg. J. Szweykowski, 18.09.1988 (POZW 487/88); Mt Łyska Skałka, above Łysa Polana glade, on the humus, alt. 1025 m, leg. J. Szweykowski, 17.08.1990 (POZW 29361, 29360); Gęsia Szyja, S slope, on the humus, Caricetum firmae, alt. 1450 m, leg. J. Szweykowski, 4.09.1992 (POZW 33058); Mt Koszysta, N slopes, the slope of the path, alt. 1315 m, leg. J. Szweykowski, 8.09.1992 (POZW 33061); the forest below Droga Oswalda Balzera road, between Potok Filipczański and Łężne, on a stone in the forest, alt. 910 m, leg. J. Szweykowski, 22.09.1988 (POZW 661/88); rocks at the bottom of Mt. Mięguszowiecki Szczyt, by Kosowinowy Piarg, humus in rock cracks, alt. 1560 m, leg. J. Szweykowski, 28.08.1990 (POZW 29369); Mt Wołoszyn (precise location not given), leg. J. Krupa, ? (WA); Dolina Roztoki valley (precise location not given), ? (WA); HIGH TATRA MTS, Slovakia: Javorová dolina valley, below Javorová polana glade, a limestone boulder by the trail, in the forest Plagiothecio-Piceetum, alt. 1240 m, 1260 m, leg. P. Górski, 21.08.2007, 18.08.2006 (POZNB 375, 370, KRAM-B); Kolová dolina valley, limestone rocks in the in the upper montane belt, 49°13.614'N, 20°11.052'E, alt. 1525 m, leg. P. Górski, 1.09.2011 (POZNB 601); Dolina Zadné Meďodoly valley, Bránka, limestone rocks by the trail, 49°14.303'N, 20°11.122'E, alt. 1205, leg. P. Górski, 22.10.2010 (POZNB 737); Dolina Kežmarskej Bielej Vody valley, SW part of Dolina Zeleneho plesa valley, above and SW from Zelené pleso Kežmarské, near the entrance to Vlka Zmrzla dolina valley, rock walls and rock fissures, mylonites, 49°12.325'N, 20°12.965'E, alt. 1665 m, leg. P. Górski, 20.11.2012 (POZNB 1337); Kôprovnica valley, upper part, mylonite rocks below Mt Krížne, 49°11.128'N, 19°57.466’E, alt. 1735 m, leg. P. Górski, 29.08.2014 (POZNB 1841, 1842); BELIANSKE TATRA MTS: Dolina Siedmich prameňov valley, 1200 m, leg. J. Váňa, 11.07.1982 (herb. Váňa); below Široké sedlo pass and Široká dolina valley, a limestone boulder by the trail, alt. 1535, leg. P. Górski, 29.08.2006 (POZNB 374, KRAM-B); Monkova dolina valley, alt. 1420 m, leg. Kulová, det. R. Šoltés sub S. aspera, 5.11.1993, rev. P. Górski (Herbarium Múzeum Tatranského národnego parku, 1/8128); Dolina siedmich prameňov valley, above the hostel Plesnivec, alt. 1200 m, leg. J. Váňa, 11.07.1982 (KRAM-B 055066); valley of the Čierny potok stream, below the rocks of Biela stena (near an unmarked route to Stará Poľana glade), calcareous rocks, 49°16.077'N, 20°11.780'E, alt. 1060 m, leg. P. Górski, 24.06.2012 (POZNB 1199, 1206); Dolina Hučava valley, middle part, calcareous rocks near the stream, 49°13.190'N, 20°18.479'E, alt. 925 m, leg. P. Górski, 19.11.2012 (POZNB 1423); Dolina pod Košiare valley, calcareous rocks below Veľký Košiar, 49°14.488'N, 20°15.040'E, alt. 1285 m, leg. P. Górski, 27.06.2012 (POZNB 1210), Dolina pod Košiare valley, calcareous rocks in the lower part of Veľký Košiar, 49°14.439'N, 20°14.948'E, alt. 1380 m, leg. P. Górski, 27.06.2012 (POZNB 1173, 1214); E part of Tristarský žľab gully, near Žľabina, 49°15.269'N, 20°12.712'E, alt. 1355 m, leg. P. Górski, 25.06.2013 (POZNB 1176, 1211, 1213); the lower part of Tristarský žľab gully, calcareous rocks in the spruce forest, 49°15.444'N, 20°12.747'E, alt. 1210 m, leg. P. Górski, 25.06.2012 (POZNB 1175); the lower part of Tristarský žľab gully, where an unmarked path crosses, 49°15.469'N, 20°12.719'E, alt. 1175 m, leg. P. Górski, 25.06.2012 (POZNB 1759); Havrania dolina valley, Havrania rovien, rocks on the threshold of the upper floor of the valley, 44°15.379'N, 20°11.301'E, alt. 1460 m, leg. P. Górski, 26.06.2012 (POZNB 1224); Havrania dolina valley, the lower part of Novy žľab gully, 49°15.951'N, 20°11.154'E, alt. 1225 m, leg. P. Górski, 26.06.2012 (POZNB 1254); the N slope of Mt. Malý vrch, the outcrops of the limestone rocks in the spruce forest, 49°15.999'N, 20°10.982'E, alt. 1205 m, leg. P. Górski, 26.06.2012 (POZNB 1193); the N slope of Mt. Malý vrch, the outcrops of the limestone rocks in the spruce forest, 49°15.999'N, 20°10.982'E, alt. 1205 m, leg. P. Górski, 26.06.2012 (POZNB 1198); Žlty žľab gully, the upper part of the gully, calacareous rocks, 49°15.595'N, 20°12.633'E, alt. 1155 m, leg. P. Górski, 19.11.2012 (POZNB 1397); a gully down Mt. Nižny Havran, above the SE part of Stará Poľana glade, the outcrops of the limestone rocks, 49°15.463'N, 20°11.935'E, alt. 1350 m, leg. P. Górski, 24.06.2012 (POZNB 1205); a gully down Mt. Nižny Havran, above the SE part of Stará Poľana glade, the outcrops of the limestone rocks, 49°15.429'N, 20°11.858'E, alt. 1440 m, leg. P. Górski, 24.06.2012 (POZNB 1221); Rígľanská dolina valley, limestone rocks near Rígľanský potok stream, 49°15.338'N, 20°13.679’E, alt. 1065 m, leg. P. Górski, 1.09.2014 (POZNB 1807).

Scapania aspera Bernet et M. Bernet Literature data WESTERN TATRA MTS, Slovakia: S slope of Mt Osobitá, alt. 1400-1500 m, leg. A. Boros, 19.08.1962 (DUDA & VÁŇA 1970a). New localities WESTERN TATRA MTS, Poland: Dolina ku Dziurze valley, near Jaskinia Dziura cave, 49°16.309'N, 19°56.575'E, alt. 1005 m, leg. P. Górski, 21.11.2011, det. J. Váňa (POZNB 847); Dolina za Bramką valley, open to visitors in the final part, limestone walls, Caricetum firmae, 49°16.337'N, 19°55.022'E, alt. 970 m, leg., det. P. Górski, 21.11.2011, conf. J. Váňa (POZNB 845); Dolina Strążyska valley, limestone rocks near the mouth of the valley, 49°16.255'N, 19°55.937'E, alt. 990 m, leg., det. P. Górski, 21.11.2011, conf. J. Váňa (POZNB 846); Żleb pod Czerwienicą gully, the E slope of Mt. Jaworzyńska Czuba, alt. 1330 m, leg. K. Buczkowska, E. Chudzińska, J. Szweykowski, 4.08.1997, det. J. Szweykowski, conf. P. Górski (POZW 38161); WESTERN TATRA MTS, Slovakia: Juráňova dolina valley, Válovec, open calcareous outcrops, 49°15.310'N, 19°46.930’E, alt. 1325 m, leg., det. P. Górski, 30.09.2013, conf. J. Váňa (POZNB 1679). Excludenda

 

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HIGH TATRA MTS, Slovakia: Kôprová dolina valley, on stones near Kôprovy potok stream, leg. J. Żebrowska (HAUSBRANDT 1949), est Scapania undulata (L.) Dumort., rev. P. Górski (WA); HIGH TATRA MTS, Poland: Mt Wołoszyn, leg. J. Krupa (HAUSBRANDT 1949), est Scapania aequiloba (Schawegr.) Dumort., rev. P. Górski (WA); BELIANSKIE TATRA MTS: precise location not given, leg. L. Goetzen-Hausbrandt (HAUSBRANDT 1949), est Scapania aequiloba (Schawegr.) Dumort., rev. P. Górski (WA); Mt Čuba, leg. V. Krajina, 23.07. 1925, det. J. Vilhelm (VILHELM 1927, PILOUS & DUDA 1960), est Plagiochila porelloides (Torrey ex Nees) Lindenb, rev. J. Duda (DUDA & VÁŇA 1970a); Červená hlina, near Alabastrová jaskyňa cave, leg. A. Boros (BOROS 1942, SZWEYKOWSKI 1958a), est Scapania aequiloba (Schawegr.) Dumort., rev. J. Duda (DUDA & VÁŇA 1970a).

Scapania brevicaulis Taylor Literature data WESTERN TATRA MTS, Poland: Dolina Jarząbcza valley, above Hala Jarząbcza Wyżnia, granite rocks above the stream, alt. 1480 m, leg. J. Szweykowski, 18.08.1956 (SZWEYKOWSKI 1960); WESTERN TATRA MTS, Slovakia: below Tomanovské sedlo pass, alt. 1575 m (ŠMARDA 1961); the N slope of Mt Poľská Tomanová, alt. 1700 m (ŠMARDA 1961); Diera, alt. 1500 m (ŠMARDA 1961); Rozpadlá dolinka valley (Rozpadliny), alt. 1690 m, 1615 m (ŠMARDA 1961, UNAR et al. 1984─1985); HIGH TATRA MTS, Poland: Szpiglasowa Przełęcz pass, mylonites on the northern side, alt. 2100 m, leg. J. Szweykowski, 27.08.1956 (SZWEYKOWSKI 1956b, 1960); HIGH TATRA MTS, Slovakia: Velická dolina valley, Mt Velická próba, alt. 1950 m, leg. J. Szweykowski, 9.07.1957 (DUDA & VÁŇA 1970b); Velická dolina valley, Mt Velická próba, alt. 1800 m (ŠMARDA 1961); Velická dolina valley, alt. 1700 m (ŠMARDA 1961); Mt Veĺká Svišťovka, leg. J. Fitzgerald (DUDA & VÁŇA 1970b); Mt Veĺká Svišťovka, alt. 1700 m, leg. J. Šmarda, 14.07.1958 (BOROS 1959, BOROS et al. 1960, ŠMARDA 1961); Mlynická dolina valley, above Skok waterfall, alt. 1825 m, leg. V. Krajina, 19.08.1932, det. J. Dostál (KRAJINA 1933); Hlinská dolina valley, alt. 1850 m (ŠMARDA 1961); Hlinská dolina valley, Malá Zahrádka, alt. 1750 m (ŠMARDA 1961); Mengusovská dolina valley, Žabie pleso lake, alt. 1700 m (ŠMARDA 1961); the W slope of Mt Kôpky, alt. 1900 m (ŠMARDA 1961); Mt Osterva, alt. 1700 m (ŠMARDA 1961); BELIANSKIE TATRA MTS: Mt Zadné Jatky, alt. 2000 m, leg. J. Váňa, 5.09.1967 (DUDA & VÁŇA 1970b); Mt Predné Jatky, alt. 1950 m, leg. J. Szweykowski, 7.07.1957 (DUDA & VÁŇA 1970b); Mt Hlúpy, alt. 2000 m, 2062 m (BOROS et al. 1960, ŠMARDA 1961); Mt Malý Košiar, alt. 1900 m (ŠMARDA 1961); Mt Kozí chrbát, alt. 1550 m (ŠMARDA 1961).

Scapania calcicola (Arnell et J. Perss.) Ingham Literature data WESTERN TATRA MTS, Poland: Mt Bobrowiec, on limestone rocks on the southern slope below Przełęcz Bobrowiecka pass, alt. 1280 m, leg. J. Szweykowski, 19.08.1956 (SZWEYKOWSKI 1960); Dolina Kościeliska valley, on limestone rocks above Kirowa Woda stream, near Hala Pisana glade, alt. 1040 m, 1065 m, leg. J. Szweykowski, 22.08.1956 (SZWEYKOWSKI 1960); BELIANSKIE TATRA MTS: Belianskie Tatra Mts, precise location not given (HAUSBRANDT 1949); Mt Ždiarska vidla, on damp limestone rocks on the eastern slope, alt. 1950 m, leg. J. Szweykowski, 19.07.1956 (SZWEYKOWSKI 1960); Mt Bujači vrch, alt. 1950 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); Mt Faixová, alt. 1480 m, 1490 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); the tourist path from the hut Protĕž until Przełęcz Pod Schody pass, on damp and shady limestone rocks below pass, alt. 1500 m, leg. J. Szweykowski, 18.07.1956 (SZWEYKOWSKI 1960); Skalné vráta pass, alt. 1600 m, leg. J. Podpěra, det. J. Duda (DUDA & VÁŇA 1968); Skalné vráta pass, alt. 1600 m, leg. J. Duda, 7.1949 (DUDA 1955a, DUDA & VÁŇA 1968); Havrania dolina valley, alt. 900 m (ŠMARDA 1961); Spod Šiky, alt. 1100 m (ŠMARDA 1961); Mt Hlúpy vrch, alt. 2000 m (ŠMARDA 1961); Mt Hlúpy vrch, alt. 2062 m, leg. J. Šmarda, 15.07.1958 (BOROS et al. 1960, DUDA & VÁŇA 1968); Mt Limbová veža, leg. J. Szweykowski (ŠMARDA et al. 1962). New localities WESTERN T ATRA M TS , Poland: Wielkie Koryciska valley, alt. 990 m, leg. P. Górski, 18.09.2005 (POZNB 365); Dolina Chochołowska valley, Jasiorów Żleb gully, above Polana Chochołowska, on shady limestone rocks, alt. 1285 m, leg. P. Górski, 8.08.2004 (POZNB 831); Polana na Stołach glade, limestone rocks amongst meadows, 49°14.977'N, 19°51.362'E, alt. 1340 m, leg. P. Górski, 20.11.2011 (POZNB 810); Dolina za Bramką valley, high, vertical rock walls, alt. 920 m, 49°16.584'N, 19°55.159'E, leg. P. Górski, 21.11.2011 (POZNB 755); Dolina ku Dziurze valley, near Jaskinia Dziura cave, open to visitors in the final part, alt. 1005 m, 49°16.318'N, 19°56.549'E, leg. P. Górski, 21.11.2011 (POZNB 758); Dolina Jaworzynka valley, Wyżnia Rówień Jaworzynka, limestone rocks, Carici-Festucetum tatrae, alt. 1100 m, leg. P. Górski, 16.09.2004 (POZNB 805); WESTERN T ATRA M TS , Slovakia: Mt Furkaska, alt. 1491 m, leg. P. Górski, 30.09.2013 (POZNB 1720); S slope of Mt Osobitá, open calcareous outcrops above Sedlo pod Osobitou pass, 49°15.561'N, 19°43.313’E, 49°15.606'N, 19°43.273'E, alt. 1555 m, leg. P. Górski, 1.10.2013 (POZNB 1682, 1686, 1690).

Scapania cuspiduligera (Nees) Müll. Frib. Literature data WESTERN TATRA MTS, Poland: Mt Kominiarski Wierch, S slope above Przełęcz Iwaniacka pass, alt. 1750 m, leg. J. Szweykowski, 13.09.1958 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, on the peak, alt. 1829 m, leg. J. Szweykowski, 20.08.1956

 

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(SZWEYKOWSKI 1960); Dolina Kościeliska valley, above Kirowa Woda steram, near Hala Pisana glade, alt. 1040 m, 1060 m, leg. J. Szweykowski, 22.08.1956 (SZWEYKOWSKI 1960); pass between Mt Małołączniak and Mt Wielka Turnia, alt. 1860 m, leg. J. Szweykowski, 11.09.1955 (SZWEYKOWSKI 1960); Dolina Jaworzynki valley, leg. J. Żebrowska (HAUSBRANDT 1949); pass between Mt Małołączniak and Mt Kopa Kondracka, alt. 1960 m, leg. J. Szweykowski, 8.09.1955 (SZWEYKOWSKI 1960); Mt Giewont, S slope below the peak, alt. 1830 m, leg. J. Szweykowski, 9.09.1955 (SZWEYKOWSKI 1960); Mt Giewont, Przełęcz Szczerba pass, humus between limestone rocks, alt. 1815 m, leg. J. Szweykowski, 9.09.1955 (SZWEYKOWSKI 1960); Dolina Strążyska valley, alt. 893-1040 m (KRUPA 1882, SZYSZYŁOWICZ 1885); WESTERN TATRA MTS, Slovakia: Tomanová dolina valley, alt. 1500 m (ŠMARDA 1961); Rozpadlá dolinka valley (Rozpadliny), alt. 1600 m (ŠMARDA 1961); Mt Rozpadlý hrebeň, alt. 1645 m (ŠMARDA 1961); HIGH TATRA MTS, Poland: Dolina Pięciu Stawów Polskich valley, mylonites, Szpiglasowa Przełęcz pass, N slope, alt. 2100 m, leg. J. Szweykowski, 27.08.1956 (SZWEYKOWSKI 1960); Dolina Pięciu Stawów Polskich valley, mylonites, Szpiglasowa Przełęcz pass, from the northern part, alt. 2120 m, leg. J. Szweykowski, 10.09.1958, as S. cuspiduligera var. tatrica (SZWEYKOWSKI 1960); HIGH TATRA MTS, Slovakia: Mt Veĺká Svišťovka, alt. 1700 m (BOROS et. al. 1960, ŠMARDA 1961, DUDA & VÁŇA 1968); Mlynická dolina valley (VÁŇA & DUDA 1965); Velická dolina valley, Mt Velická próba, alt. 1950 m, leg. J. Szweykowski, 9.07.1957 (DUDA & VÁŇA 1968); Velická dolina valley, alt. 1700 m (ŠMARDA 1961); Kôprovské sedlo pass, alt. 2040 m (ŠMARDA 1961); W slope of Mt Kôpky, alt. 1900 m (ŠMARDA 1961); Mt Ostrva, alt. 1700 m (ŠMARDA 1961); Javorová dolina valley (ŠMARDA 1961); BELIANSKIE TATRA MTS: Mt Ždiarska vidla, alt. 2148 m (ŠMARDA 1938); Mt Ždiarska vidla, alt. 1830 m, leg. J. Váňa, 9.09.1967 (DUDA & VÁŇA 1968); Mt Ždiarska vidla, N slope below the peak, alt. 2100─2130 m, leg. J. Szweykowski, 19.07.1956 (SZWEYKOWSKI 1960); Mt Ždiarska vidla, E slope, near the route, alt. 1950 m, leg. J. Szweykowski, 19.07.1956 (SZWEYKOWSKI 1960); Mt Zadné Jatky, below the summit, alt. 2000 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Zadné Jatky, N slope, snow-beds, alt. 1800 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Predné Jatky, on the humus soil on the pass, alt. 1900 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Predné Jatky, on the soil between limestone rocks, on the second peak from the south, alt. 1980 m, leg. J. Szweykowski, 22.07. 1956 (SZWEYKOWSKI 1960); Mt Bujači vrch, Caricetum firmae, between limestone rocks, on the northern slope, alt. 1710 m, leg. J. Szweykowski, 18.07.1956 (SZWEYKOWSKI 1960); the ridge between Mt Bujači vrch and Mt Predné Jatky, on the soil in rock cracks, alt. 1900 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Bujači vrch, the NW slope, alt. 1850 m, leg. J. Szweykowski, 18.07.956 (SZWEYKOWSKI 1960); Mt Bujači vrch, on the humus, on the peak, alt. 1950 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); Mt Kozi chrbát, on a damp, limestone rock, by a tourist trail, on the northern side, eastwards of Skalné vráta pass, alt. 1600 m, leg. J. Szweykowski, 20.07.1956 (SZWEYKOWSKI 1960); Skalné vráta pass, the cracks of limestone rocks, alt. 1620 m, leg. J. Szweykowski, 18.07.1956 (SZWEYKOWSKI 1960); the route between Sedlo Pod Schody pass and Skalné vráta pass, on the slope, alt. 1600 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); Mt Kozi chrbát, on the soil between limestone rocks, on the northern slope between Mt Faixová and Skalné vráta pass, alt. 1500 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); the path from the hut Protez to Sedlo Pod Schody pass, on shady limestone rocks, under the pass, alt. 1500 m, leg. J. Szweykowski, 18.07.1956 (SZWEYKOWSKI 1960); Mt Hlúpy vrch, alt. 2062 m (ŠMARDA 1938, BOROS et. al. 1960); Mt Hlúpy vrch, alt. 2050 m, leg. J. Váňa, 5.09.1967 (DUDA & VÁŇA 1968); Mt Zadné Jatky, alt. 2024 m (ŠMARDA 1938, BOROS et. al. 1960, BOROS 1959; DUDA & VÁŇA 1968); Tristarská dolina valley, between Mt Ždiarska vidla and Mt Havran, leg. Z. Pilous, det. J. Duda (DUDA & VÁŇA 1968); between Mt Hlúpy vrch and Mt Zadné Jatky, alt. 2000 m (ŠMARDA 1938); between Mt Hlúpy vrch and Mt Zadné Jatky, alt. 1900 m, leg. J. Váňa, 6.09.1967 (DUDA & VÁŇA 1968); Mt Malý Košiar, between Mt Predné and Mt Zadné Jatky, alt. 1920 m, leg. J. Váňa, 6.09.1967 (DUDA & VÁŇA 1968); Mt Predné Jatky, alt. 2000 m, leg. J. Váňa, 6.09.1967 (DUDA & VÁŇA 1968); Mt Bujači vrch, alt. 1950 m, leg. A. Vězda, 16.08.1947 (DUDA 1958, DUDA & VÁŇA 1968); Mt Bujači vrch, alt. 1930 m, leg. J. Váňa, 6.09.1967 (DUDA & VÁŇA 1968); Skalné vráta pass, alt. 1600 m, leg. J. Podpěra, 21.07.1922, det. J. Duda (DUDA & VÁŇA 1968); Skalné vráta pass, alt. 1623 m, leg. A. Vežda, 15.08.1947 (DUDA 1958, DUDA & VÁŇA 1968); Dolina siedmich prameňov valley, leg. J. Růžička, 26.08.1957 (DUDA & VÁŇA 1968); Faixová lúka, alt. 1500-1600 m, leg. A. Boros, 11.09.1940 (DUDA & VÁŇA 1968); Havrania dolina valley (Dominův důl), between Mt Havran and Mt Nový, alt. 1750 m (ŠMARDA 1938); Havrania dolina valley, alt. 1700 m, leg. J. Suza, det. J. Šmarda (SUZA 1937, DUDA & VÁŇA 1968); Mt Havran, alt. 2154 m (ŠMARDA 1938); Mt Havran, alt. 2000─2150 m (SUZA 1937); Široké pole, alt. 1700 m (ŠMARDA 1938); Košiary (between Mt Zadné and Mt Predné Jatky), alt. 2011 m (ŠMARDA 1938); Mt Bujači vrch, alt. 1800 m (ŠMARDA 1956); Skalné vráta pass, alt. 1620 (ŠMARDA et al. 1962); Mt Rakúsky chrbát (ŠMARDA et al. 1962); Kozí chrbát, Mt Jelenia skala (ŠMARDA et al. 1962). New localities HIGH TATRA MTS, Poland: Mięguszowiecka Przełęcz pod Chłopkiem pass, rock crevices, 49°11.018'N, 20°03.920’E, alt. 2307 m, leg. P. Górski, 2.09.2014 (POZNB 1889); Dolina Rybiego Potoku valley, near Morskie Oko lake, alt. 1393 m, leg. F. Lilienfeldówna, T. Wilczyński, 29.08.1912, det. P. Górski (S. cuspiduligera was found as admixture in a voucher specimen named as Gymnomitrion adustum and stored in WA); HIGH TATRA MTS, Slovakia: Dolina Kežmarskej Bielej Vody valley, SW part of Dolina Zeleneho plesa valley, above and SW from Zelené pleso Kežmarské, near the entrance to Vlka Zmrzla dolina valley, rock walls and rock fissures, mylonites, 49°12.325'N, 20°12.965'E, alt. 1665 m, leg. P. Górski, 20.11.2012 (POZNB 1336); Kôprovnica valley, upper part, rocks below Mt Krížne, 49°11.157'N, 19°57.485’E, alt. 1740 m, leg. P. Górski, 29.08.2014 (POZNB 1845); WESTERN T ATRA M TS , Poland: Kobylarzowy Żleb pod Małołączniakiem gully, in the upper part, rock walls N exposed, alpine grassland with Carex firma, on the humus or bare rock wall, 49°14.632'N, 19°54.649'E, alt. 1700 m, leg. P. Górski, 22.11.2011 (POZNB 781); Kobylarzowy Żleb pod Małołączniakiem gully, in the upper part, limestone rocks N exposed, 49°14.668'N, 19°54.757'E, alt. 1760 m, leg. P. Górski, 22.11.2011 (POZNB 787); Wyżnia Kondracka Przełęcz pod Giewontem pass, alpine grassland Festuco-Seslerietum tatrae, alt. 1785 m, leg. P. Górski, 11.09.2004 (POZNB 381); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), in the lower part of the wetlands, springfens with Cratoneuron, 49°15.107'N, 19°50.972'E, alt. 1180 m, leg. P. Górski, 20.11.2011 (POZNB 817); Żleb Zabijak gully (the upper part of the Dolina Lejowa valley), in the middle part, the limestone rock with Ctenidium, 49°15.053'N, 19°50.945'E, alt. 1215 m, leg. P. Górski, 20.11.2011 (POZNB 796); Polana na Stołach glade, limestone rocks amongst meadows, 49°14.977'N, 19°51.362'E, alt. 1340 m, leg. P. Górski, 20.11.2011 (POZNB 770); Skupniów Upłaz, near the end of the trail down to

 

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Dolina Jaworzynki valley, alt. 1450 m, leg. P. Górski, 16.09.2004, (POZNB 380); Dolina Jaworzynka valley, Wyżnia Rówień Jaworzynka, limestone rocks Carici-Festucetum tatrae, alt. 1100 m, leg. P. Górski, 16.09.2004 (POZNB 814); W ESTERN T ATRA MTS , Slovakia: Mt Sivý vrch, alt. 1600 m, leg. V. Peciar, 28.09.1976 (herb. Váňa; dupl. ex SLO); BELIANSKIE TATRA MTS: Mt Faixová, alt. 1480 m, leg. J. Váňa, 1.09.1972 (herb. Váňa); Skalné vráta pass, alt. 1600 m, leg. J. Váňa, 10.07.1982 (herb. Váňa); Mt Ždiarska vidla, northern slopes in the sub-peak part, alt. 1980 m, 2040 m, leg. P. Górski, 19.08.2006 (POZNB 382, 815); Dolina pod Košiare valley, a little above the place linking the stream from Jahňacia dolina valley and Tokársky potok stream, calcareous rocks, 49°14.894'N, 20°15.548'E, alt. 1030 m, leg. P. Górski, 27.06.2012 (POZNB 1174); the gully down the Mt Nižny Havran, above the SE part of Stará Poľana glade, the outcrops of limestone rocks, 49°15.463'N, 20°11'935'E, alt. 1350 m, leg. P. Górski, 24.06.2012 (POZNB 1204); Mt Havran, on the peak, 49°14.910'N, 20°11.857'E, alt. 2154 m, leg. P. Górski, 26.06.2012 (POZNB 1208); the E part of Tristarský žľab gully, near Žľabina, 49°15.269'N, 20°12.712'E, alt. 1355 m, leg. P. Górski, 25.06.2013 (POZNB 1212); Dolina pod Košiare valley, calcareous rocks in the lower part of Veľký Košiar, 49°14.439'N, 20°14.948'E, alt. 1380 m, leg. P. Górski, 27.06.2012 (POZNB 1217); Havrania dolina valley, Havrania rovien, rocks on the treshold of the upper floor of the valley, 44°15.379'N, 20°11.301'E, alt. 1460 m, leg. P. Górski, 26.06.2012 (POZNB 1223); Havrania dolina valley, Havrania rovien, rocks on the treshold of the upper floor of the valley, 44°15.379'N, 20.11.301'E, alt. 1460 m, leg. P. Górski, 26.06.2012 (POZNB 1281); Dolina pod Košiare valley, calcareous rocks in the lower part of Veľký Košiar, 49°14.439'N, 20°14.948'E, alt. 1380 m, leg. P. Górski, 27.06.2012 (POZNB 1287); Havrania dolina valley, Havrania rovien, rocks on the treshold of the upper floor of the valley, 44°15.325'N, 20°11.366'E, alt. 1550 m, leg. P. Górski, 26.06.2012 (POZNB 1289); Mt Havran, on the peak, 49°14.910'N, 20°11.857'E, alt. 2154 m, leg. P. Górski, 26.06.2012 (POZNB 1294).

Scapania gymnostomophila Kaal. Literature data WESTERN TATRA MTS, Poland: Mt Kominiarski Wierch, humus between limestone rock, S slope, above Przełęcz Iwaniacka pass, alt. 1500 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, humus between limestone rocks, on the southern slope above Przełęcz Iwaniacka pass, alt. 1745 m, leg. J. Szweykowski, 13.09.1958 (SZWEYKOWSKI 1956a, 1958b, 1960); Kalackie Koryto (Suchy Żleb) gully, N slope of Mt Kalacka Turnia, alt. 1300 m, leg. H. Klama (SZWEYKOWSKI & KLAMA 2010; herb. H. Klama); Mt Stoły, SSW slope, above Tomanowa Przełęcz pass, alt. 1775 m, leg. H. Klama (SZWEYKOWSKI & KLAMA 2010; herb. H. Klama); WESTERN TATRA MTS, Slovakia: Mt Sivý vrch, alt. 1600 m (PECIAR 1981); BELIANSKIE TATRA MTS: Mt Ždiarska vidla, humus between rocks on the northern slope below the peak, alt. 2100─2130 m, 19.07.1956 (SZWEYKOWSKI 1958b, 1960); between Mt Bujači vrch and Mt Predné Jatky, in the cracks of limestone rock, alt. 1900 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1958b, 1960, ŠMARDA 1961); Mt Bujači vrch, humus between limestone rocks on the NE side, alt. 1710 m, leg. J. Szweykowski, 18.07.1956 (SZWEYKOWSKI 1960); Mt Bujači vrch, alt. 1930 m, leg. J. Váňa, 6.09.1967 (DUDA & VÁŇA 1968); Mt Faixová, humus between limestone rocks on the peak, alt. 1490 m, 21.07.1956 (SZWEYKOWSKI 1958b, 1960, ŠMARDA 1961); Mt Hlúpy, alt. 2000 m (ŠMARDA 1961); Mt Hlúpy, on the peak, alt. 2062 m (BOROS et al. 1960); Mt Hlúpy, SW slope (BOROS et al. 1960); Mt Hlúpy, N slope (BOROS et al. 1960); Mt Hlúpy, alt. 2050 m, leg. J. Váňa, 1967 (DUDA & VÁŇA 1968); Mt Predné Jatky, alt. 1950 m, leg. J. Szweykowski, 7.07.1957 (DUDA & VÁŇA 1968); Mt Veĺká Svišťovka, SW slope, alt. 1700 m (BOROS et al. 1960, ŠMARDA 1961); Mt Predné Jatky, N slope, alt. 1950 m, leg. J. Szweykowski (DUDA & VÁŇA 1968); Mt Zadné Jatky, Kamzičia jaskýna cave, alt. 1900 m (ŠMARDA 1961); Mt Zadné Jatky, alt. 2024 m (BOROS 1959). New localities WESTERN T ATRA M TS , Poland: Dolina Kościeliska valley, southwards of Polana Pisana glade, limestone rocks by the trail to the hostel on Hala Ornak glade, alt. 1035 m, leg. P. Górski, 27.08.2008 (POZNB 371); Dolina Lejowa valley, limestone rocks at mouth of the valley, 49°16.181'N, 19°50.537'E, alt. 965 m, leg. P. Górski, 20.11.2011 (POZNB 771); the upper part of the Kobylarzowy Żleb pod Małołączniakiem gully, limestone rocks N exposed, 49°14.668'N, 19°54.757'E, alt. 1760 m, leg. P. Górski, 22.11.2011 (POZNB 848); Kalackie Koryto (Suchy Żleb) gully, in the middle part of the gully, on a limestone boulder N exposed, 49°15.357'N, 19°57.610'E, alt. 1335 m, leg. P. Górski, 24.11.2011 (POZNB 775; from Kalackie Koryto gully this species was recorded earlier by H. Klama, see Literature data); WESTERN T ATRA M TS , Slovakia: Mt Sivý vrch, alt. 1600 m, leg. V. Peciar, 28.09.1976 (herb. Váňa, dupl. ex SLO); Mt Osobitá, on the paek, alt. 1687 m, leg. P. Górski, 1.10.2013 (POZNB 1698); S slope of Mt Osobitá, open calcareous outcrops above Sedlo pod Osobitou pass, 49°15.561'N, 19°43.313’E, 49°15.606'N, 19°43.273'E, alt. 1555 m, 1610 m, leg. P. Górski, 1.10.2013 (POZNB 1683, 1687, 1688); HIGH TATRA MTS, Poland: below Mięguszowiecka Przełęcz pod Chłopkiem pass, rock crevices, 49°11.037'N, 20°03.955’E, 49°11.018'N, 20°03.920’E, alt. 2305 m, 2307 m, leg. P. Górski, 2.09.2014 (POZNB 1909, 1891); Mt Kazalnica Cubryńska near Morskie Oko lake, climbing route "Jagiełło-Roj", rock fissures, 49°11'32''N, 20°03'47''E, alt. 1700 m, leg. S. Szlagowski, 15.03.2012, det. P. Górski, conf. J. Váňa (POZNB 1017; see also GÓRSKI & VÁŇA in ELLIS et. al. 2013); HIGH TATRA MTS, Slovakia: Dolina Kežmarskej Bielej Vody valley, the SW part of Dolina Zeleneho plesa valley, above and SW from Zelené pleso Kežmarské, near the entrance to Vlka Zmrzla dolina valley, rock walls and rock fissures, mylonites, 49°12.325'N, 20°12.965'E, alt. 1665 m, leg. P. Górski, 20.11.2012 (POZNB 1342); Kôprovnica valley, upper part, rocks below Mt Krížne, 49°11.128'N, 19°57.466’E, alt. 1735 m, leg. P. Górski, 29.08.2014 (POZNB 1841); BELIANSKIE TATRA MTS: Mt Havran, on the peak, 49°14.910'N, 20°11.857'E, alt. 2154 m, leg. P. Górski, 26.06.2012 (POZNB 1209); Mt Faixová, alt. 1480 m, leg. J. Váňa, 1.09.1972 (herb. Váňa); Skalné vráta pass, Pod Schody, alt. 1500 m, leg. J. Váňa, 2.09.1976 (herb. Váňa).

 

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Scapania helvetica Gottsche Literature data WESTERN TATRA MTS, Poland: Dolina Kondratowa valley, Polana Kalatówki glade, alt. 1170 m, leg. F. Lilienfeldówna, 27.08.1912 (LILIENFELDÓWNA 1914b); Wyżnia Dolina Chochołowska valley, on a granite waste, alt. 1445 m, 1480 m, leg. J. Szweykowski, 26.07.1957 (SZWEYKOWSKI 1960); Dolina Chochołowska valley, Litworowy Żleb gully, on wet soil, alt. 1340 m, leg. J. Szweykowski, 31.071957 (SZWEYKOWSKI 1960); Dolina Chochołowska valley, on soil, by the estuary of Dolina Starorobociańska valley, alt. 1100 m, leg. J. Szweykowski, 11.09.1958 (SZWEYKOWSKI 1960); Dolina Jarząbcza valley, Jarząbcza Rówień, on a granite boulder by the route, alt. 1480 m, leg. J. Szweykowski, 29.07.1957 (SZWEYKOWSKI 1960); Dolina Jarząbcza valley, W slope of Mt Czubik, on wet soil near stream, alt. 1495 m, leg. J. Szweykowski, 31.07.1957 (SZWEYKOWSKI 1960); Dolina Jarząbcza valley, between Mt Czubik and Mt Jarząbczy Wierch, alt. 1445 m, leg. J. Szweykowski, 20.07.1957 (SZWEYKOWSKI 1960); Mt Jarząbczy Wierch, W slope, above Dolina Jarząbcza valley, alt. 1685 m, leg. J. Szweykowski, 5.08.1956 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, on the paek, alt. 1829 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, E slope, above Zdrapiska, alt. 1750 m, leg. J. Szweykowski, 20.08.1956 (SZWEYKOWSKI 1958c, 1960); the pass between Mt Małołączniak and Mt Wielka Turnia, alt. 1860 m, leg. J. Szweykowski, 11.09.1955 (SZWEYKOWSKI 1960); Mt Kominiarski Wierch, S slope above Przełęcz Iwaniacka pass, alt. 1750 m, leg. J. Szweykowski, 13.09.1958 (SZWEYKOWSKI 1958c, 1960; POZW, WA); the pass between Mt Małołączniak and Mt Kopa Kondracka, humus between limestone rocks, alt. 1960 m, leg. J. Szweykowski, 8.09.1955 (SZWEYKOWSKI 1960); Mt Kopa Kondracka, N slope, alt. 1920 m, leg. J. Szweykowski, 26.06.1954 (SZWEYKOWSKI 1960); Mt Giewont, Przełęcz Szczerba pass, humus between limestone rocks, alt. 1815 m, 9.09.1955 (SZWEYKOWSKI 1960); Dolina Olczyska valley, near the route, alt. 980 m, leg. J. Szweykowski, 19.07.1948; WESTERN TATRA MTS, Slovakia: Mt Plačlivý Roháč, S slope, alt. 2000 m, leg. J. Váňa, 12.09.1966 (DUDA & VÁŇA 1969a); Mt Velký Baranec, alt. 2187 m (DUDA 1955a); Mt Velký Baranec, the NE slope, alt. 2000 m, leg. J. Duda, 13.09.1966 (DUDA & VÁŇA 1969a); Trnovská dolina valley, alt. 900 m (ŠMARDA 1961); Bystrá dolina valley, near the Bystrá stream, alt. 1000 m (ŠMARDA 1961); Bystrá dolina valley, Bystré plesa lakes, alt. 2000 m (ŠMARDA 1961); Mt Temniak, alt. 2100 m (ŠMARDA 1961); Rozpadlá dolinka valley (Rozpadliny), alt. 1690 m (ŠMARDA 1961); Tomanová dolina valley, alt. 1500 m (ŠMARDA 1961); Mt Polská Tomanová, alt. 1700 m (ŠMARDA 1961); HIGH TATRA MTS, Poland: Morskie Oko lake, leg. T. Wiśniewski (HAUSBRANDT 1949; WA); screes above Morskie Oko lake, Oxyrio-Saxifragetum carpathicae, alt. 1500 m (PAWŁOWSKI et al. 1928); Dolina Gąsienicowa valley, sub-glacially eroded step of Kozia dolinka valley, on granite rocks, alt. 1835 m, leg. J. Szweykowski, 3.08.1957 (SZWEYKOWSKI 1960); glacial cirque of Zmarzły Staw Gąsienicowy lake, below Zawrat pass, on mylonite wastes, alt. 2020 m, leg. J. Szweykowski, 3.08.1957 (SZWEYKOWSKI 1960); Mt Mały Kozi Wierch, S slope, above Dolinka Pod Kołem valley, on granite rocks, alt. 2050 m, leg. J. Szweykowski, 26.08.1956 (SZWEYKOWSKI 1960); Dolina Pięciu Stawów Polskich valley, mylonites on the E slope of Niżni Kostur, alt. 1800 m, leg. J. Szweykowski, 9.09.1958 (Szweykowski 1960); Dolina Pięciu Stawów Polskich valley, E slope of Mt Wyżni Kostur, mylonites, alt. 1950 m, leg. J. Szweykowski, 9.09.1958 (SZWEYKOWSKI 1960); Dolina za Mnichem valley, on soil near stream, alt. 1830 m, leg. J. Szweykowski, 10.09.1958 (SZWEYKOWSKI 1960); Dolina Pięciu Stawów Polskich valley, Minuartio-Oreochloetum distichae, alt. 2200 m (BALCERKIEWICZ 1984); Sołtysówka stream valley, wet soil on path in the spruce forest, alt. 1000 m (KLAMA 2008); HIGH TATRA MTS, Slovakia: Mt Lomnický štít, alt. 2633 m, leg. I. Szyszyłowicz (SZYSZYŁOWICZ 1885, KULESZA 1920, HAUSBRANDT 1949); Veĺka Studená dolina valley, near Vareškové pleso lake, alt. 1833 m, leg. I. Györrfy, 5.08.1910 (SCHIFFNER 1911); Mlynická dolina valley, above Skok waterfall, near stream, Hygrohypnetum dilatati, alt. 1885, leg. V. Krajina, 21.08.1932, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, Mt Štrbské Solisko, Oxyrio-Saxifragetum carpathicae, alt. 2100 m, leg. V. Krajina, 9.09.1931, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, above Capie pleso lake, below Bystré sedlo pass, Luzuletum alpino-pilosae, alt. 2090 m, leg. V. Krajina, 1.09.1931, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, below Skok waterfall, Calthetum laetae, alt. 1650 m, 1740 m, leg. V. Krajina, 31.07.1932, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, near Nižne Kozie pleso lake, Aconitetum firmi, alt. 1955 m, leg. V. Krajina, 4.09.1931, det. J. Dostál (KRAJINA 1933, VÁŇA & DUDA 1965); Mlynická dolina valley, above Skok waterfall, Trisetetum fusci, alt. 1840 m, leg. V. Krajina, 21.08.1932, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, above Skok waterfall, Salicetum lapponum, alt. 1815 m, 1830 m, leg. V. Krajina, 5.09.1931, 19.08.1932, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, above Skok waterfall, Drepanocladetum exannulati, alt. 1840 m, leg. V. Krajina, 19.08.1932, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, Pleso nad Skokom lake, leg. J. Szweykowski, 16.07.1958 (DUDA & VÁŇA 1969a); Mlynická dolina valley, near Pleso nad Skokom lake, alt. 1810 m (ŠMARDA 1961); Mlynická dolina valley, below Nižne Kozie pleso lake, Drepanocladetum exannulati, alt. 1890 m, 1895 m, leg. V. Krajina, 10.08.1932, det. J. Dostál (KRAJINA 1933); Mlynická dolina valley, above Mlynica stream, alt. 1600 m, leg. J. Szweykowski, 15.07.1958 (DUDA & VÁŇA 1969a); Mt Velké Solisko, alt. 2260 m, leg. V. Krajina, 4.08.1931, det. J. Váňa, J. Duda (VÁŇA & DUDA 1965); Mlynická dolina valley, on wet soil near stream, alt. 1700 m, leg. J. Szweykowski, 16.08.1958 (SZWEYKOWSKI 1960); Dolina Kežmarskej Bielej Vody valley, on wet soil near Čierne pleso lake, alt. 1580 m, leg. J. Szweykowski, 23.07.1956 (SZWEYKOWSKI 1960); Važecká dolina valley, near Krivánske Zelené pleso lake, leg. J. Růžička, det. J. Duda sub S. irrigua, rev. J. Duda (DUDA 1962, DUDA & VÁŇA 1969a); near Ľadový potok stream, alt. 1700 m, leg. J. Jedlička, 21.09.1950, det. J. Duda (DUDA & VÁŇA 1969a); Batizovská dolina valley, near Batizovské pleso lake, alt. 1900 m (ŠMARDA 1961); Batizovská dolina valley, near Batizovské pleso lake, alt. 1800─1900 m, leg. J. Podpěra, 25.07.1921 (DUDA & VÁŇA 1969a); Mt Veĺká Svišťovka, alt. 1700 m, leg. J. Szweykowski, 11.07.1957 (DUDA & VÁŇA 1969a); Mt Veĺká Svišťovka, alt. 1700 m, leg. J. Šmarda, 14.07.1958 (BOROS et al. 1960, ŠMARDA 1961, DUDA & VÁŇA 1969a); Mt Veĺká Svišťovka, alt. 1800─2000 m, leg. A. Boros, 14.07.1958 (BOROS 1959, BOROS et al. 1960); Kolová dolina valley, near Kolové pleso lake, alt. 1560 m, leg. J. Růžička (DUDA 1962, DUDA & VÁŇA 1969a); Furkotská dolina valley, alt. 1900 m (ŠMARDA 1961); Bystré sedlo pass, alt. 2100 m (ŠMARDA 1961); Kôprovské sedlo pass, alt. 2140 m, 2150 m (ŠMARDA 1960a, 1961); Mengusovská dolina valley, alt. 1800 m (ŠMARDA 1961); Mengusovské sedlo pass, 2100─2300 m (ŠMARDA 1960a, 1961); below Mt Kôpky (ŠMARDA 1961); Mengusovská dolina valley, Žabie pleso lake (ŠMARDA 1961); Červený žlab gully, alt. 1610 m (ŠMARDA 1961); Hlinská dolina valley, alt. 1850 m (ŠMARDA 1961); Hlinská dolina valley, Malá Zahrádka, alt. 1750 m (ŠMARDA 1961); Zadná Tichá dolina valley, alt. 1800 m (ŠMARDA 1961); Zlomisková dolina valley, alt. 1800 m (ŠMARDA

 

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1961); Mt Ostrva, alt. 1950 m (ŠMARDA 1961); Velická dolina valley, Mt Velická próba, alt. 1800 m (ŠMARDA 1961); Dolina Kežmarskej Bielej Vody valley, Červená dolina valley, Festucetum pictae, alt. 2001 m, leg. R. Šoltés, 20.08.1993 (ŠOLTÉS et al. 2006); Dolina Kežmarskej Bielej Vody valley, Dolina Zeleného plesa valley, Oxyrio-Saxifragetum carpathicae, alt. 1691 m, leg. R. Šoltés, 20.08.2003 (ŠOLTÉS et al. 2006); Dolina Kežmarskej Bielej Vody valley, Malá Zmrzlá dolina valley, Andreaetum nivalis, alt. 1937 m, leg. R. Šoltés, 24.07.2004 (ŠOLTÉS et al. 2006); BELIANSKIE TATRA MTS: Belianskie Tatra, leg. Wiśniewski (HAUSBRANDT 1949); Mt Ždiarska vidla, N slope, the humus between limestone rocks, alt. 2100 m, leg. J. Szweykowski, 19.07.1956 (SZWEYKOWSKI 1960); Mt Predné Jatky, the E slope, alt. 1900 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Zadné Jatky, on the peak, alt. 2000 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Kozi chrbát, N slope between Sedlo pod Schody pass and Skalné vráta pass, the humus between limestone rocks, alt. 1500 m, leg. J. Szweykowski, 21.07.1956 (SZWEYKOWSKI 1960); Mt Kozi chrbát, on the slope of a tourist route between Faixová and Sedlo pod Schody pass, alt. 1500 m, leg. J. Szweykowski, 22.07.1956 (SZWEYKOWSKI 1960); Mt Hlúpy, alt. 1800─1900 m, leg. A. Boros, 15.07.1958 (DUDA & VÁŇA 1969a); Mt Hlúpy (BOROS et al. 1960); Mt Hlúpy, SW slope (BOROS et al. 1960); Mt Hlúpy, alt. 2062 m (BOROS et al. 1960); Mt Hlúpy, alt. 1900 m (ŠMARDA 1961); Mt Hlúpy, alt. 2050 m, leg. J. Váňa, 5.09.1967 (DUDA & VÁŇA 1969a); the pass between Mt Hlúpy and Mt Jatky, alt. 1900 m, leg. A. Boros, 15.07.1958 (DUDA & VÁŇA 1969a); Havrania dolina valley, 1600 m (ŠMARDA 1961); Mt Zadné Jatky, alt. 1900 m (ŠMARDA 1961); Mt Zadné Jatky, alt. 2000 m, leg. J. Szweykowski, 7.07.1957 (DUDA & VÁŇA 1969a); Mt Zadné Jatky, leg. J. Váňa, 5.09.1967 (DUDA & VÁŇA 1969a); Mt Jatky, leg. F. Hazslinszky, det. J. Duda (DUDA & VÁŇA 1969a); Mt Bujači vrch, alt. 1930 m, leg. J. Váňa, 6.09.1967 (DUDA & VÁŇA 1969a); Malý Košiar, alt. 1900 m (ŠMARDA 1961); Mt Jelenia skala, alt. 1600 m (ŠMARDA 1961; ŠMARDA et al. 1962); Široké pole, alt. 1800 m (ŠMARDA 1961); Predné Meďodoly valley, alt. 1600 m (ŠMARDA 1961); Dolina siedmich prameňov valley, alt. 1240 m, leg. I. Györrfy, 12.07.1912 (DUDA & VÁŇA 1969a). New localities HIGH TATRA MTS, Slovakia: Žabia (Mengusovská) dolina valley, Malé Žabie pleso lake, alt. 1920 m, leg. J. Váňa, 3.09.1973 (herb. Váňa); Mengusovská dolina valley, Mengusovské sedlo pass, alt. 2180 m, leg. J. Váňa, 5.09.1973 (herb. Váňa); W slope of Mt Ostrva, alt. 1700 m, leg. J. Váňa, 2.09.1973 (herb. Váňa); Velická dolina valley, Kvetnicové pleso lake, alt. 1810 m, leg. J. Váňa, 2.09.1972 (herb. Váňa); Malá Studená dolina valley, Prostredné Spišské pleso lake, alt. 2015 m, leg. J. Váňa, 3.09.1972 (herb. Váňa); Velká Studená dolina valley, Zbojnické pleso lake, alt. 1970 m, leg. J. Váňa, 30.08.1972 (herb. Váňa); Velká Studená dolina valley, Žltá stena, near Terryho chata, alt. 1950 m, leg. J. Váňa, 31.8.1972 (herb. Váňa); Červena dolina valley, near Červené pleso pond, alt. 1815 m, leg. P. Górski, 25.09.2007, conf. J. Váňa (POZNB 1057); Važecká dolina valley, the lower glacial cirque, alt. 1880 m, leg. P. Górski, 11.08.2010, conf. J. Váňa (POZNB 1064); Važecká dolina valley, near Krivánske Zelené pleso lake, Salicetum herbaceae, 49°09.455'N, 20°00.396'E, alt. 2000 m, leg. P. Górski, 11.08.2010, conf. J. Váňa (POZNB 1075); Veĺka Studená dolina valley, near the route to the mountain hostel, wet slope trail, alt. 1680 m, leg. P. Górski, 24.08.2007, conf. J. Váňa (POZNB 1087); Nefcerka valley, above E shore of Nižné Terianske pleso lake, rock walls with dripping water, 49°10.125' E 20°00.833', alt. 1960 m, leg. P. Górski, 2.08.2011, conf. J. Váňa (POZNB 1088); Nefcerka valley, above E shore of Nižné Terianske pleso lake, on peat, on stones with running water to lake, 49°10.102'N, 20°00.878', alt. 1970 m, leg. P. Górski, 2.08.2012 (POZNB 1313, 1330); Zlomisková dolina valley, Kotlina Ľodového plesa, near SE shore of Ľodové pleso lake, 49°09.771'N, 20°06.453'E, alt. 1930 m, leg. P. Górski, 6.08.2010 (POZNB 1514); Zlomisková dolina valley, the lower glacial cirque, 49°09.622'N, 20°05.981'E, alt. 1810 m, leg. P. Górski, 7.08.2012 (POZNB 1583); Hlinská dolina valley, near the route, Salicetum herbaceae, alt. 1790 m, leg. P. Górski, 21.08.2006 (POZNB 1566); Kolová dolina valley, SEE from and above Kolové pleso lake, below rock walls of Hrebeň Kolových veži, 49°13.106'N, 20°11.663'E, alt. 1645 m, leg. P. Górski, 28.08.2010 (POZNB 1760); Kôprovnica valley, upper part, rocks below Mt Krížne, 49°11.157'N, 19°57.485’E, alt. 1755 m, leg. P. Górski, 29.08.2014 (POZNB 1931); HIGH TATRA MTS, Poland: Dolina Pięciu Stawów Polskich valley, in the stream coming from Zadni Staw Polski lake, on a stone and humus, alt. 1860 m, leg., det. P. Górski, 28.07.2005, conf. J. Váňa (POZNB 1067); Dolina Pięciu Stawów Polskich valley, peat-bog above SE shore of Wielki Staw Polski, below Szpiglasowa Przełęcz pass, 49°12.155'N, 20°02.263'E, alt. 1790 m, leg. P. Górski, 24.08.2012 (POZNB 1756); Dolina Morskiego Oka valley, near the route to Morskie Oko lake, above Wodogrzmoty Mickiewicza waterfall, an earthy scarp, leg., det. P. Górski, 15.09.2004, conf. J. Váňa (POZNB 1077); WESTERN TATRA MTS, Slovakia: Spálená dolina valley, Arabido-Cratoneuretum, alt. 1735m, 1740 m, leg. P. Górski, 19.07.2006, conf. J. Váňa (POZNB 1060, 1063); N from Chata pod Náružím mountain hostel, near the route to Predúvratie pass, alt. 1470 m, leg. P. Górski, 19.07.2008 (POZNB 802); WESTERN TATRA MTS, Poland: Kondratowa dolina valley, above Polana Kondratowa glade, slope trail to Mt Giewont, alt. 1500 m, leg. P. Górski, 11.09.2004 (POZNB 803); Dolina Wyżnia Jarząbcza valley, near Mt Jarząbczy Kopieniec, on soil, alt. 1700 m, leg., det. P. Górski, 28.09.2003, conf. J. Váňa (POZNB 1054); WESTERN TATRA MTS, Slovakia: the upper part of the Kamienistá dolina valley, below Pyšne sedlo pass, near the stream, 49°11.437'N, 19°51.531'E, alt. 1680 m, leg. P. Górski, 4.08.2012 (POZNB 1757); BELIANSKIE TATRA MTS: Mt Faixová, alt. 1480 m, leg. J. Váňa, 1.09.1972 (herb. Váňa); alpine grasslands below Havranie sedlo pass, N slope, 49°15.018'N, 20°11.482'E, alt. 1805 m, leg. P. Górski, 26.06.2012 (POZNB 1584). Excludenda HIGH TATRA MTS, Poland: Mt Wołoszyn, leg. J. Krupa (HAUSBRANDT 1949), est Scapania undulata (L.) Dumort., rev. P. Górski (WA). Scapania praetervisa Meyl. Literature data WESTERN T ATRA M TS , Poland: Mt Jarząbczy Wierch, W slope, above Dolina Jarząbcza valley, alt. 1685 m, leg. J. Szweykowski, 18.08.1956 (SZWEYKOWSKI 1960).

 

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New localities WESTERN T ATRA M TS, Slovakia: Červené vrchy, Javorový žľab gully, alt. 1700 m, leg. J. Váňa, 18.08.1995 (herb. Váňa); BELIANSKIE TATRA MTS: Mt Faixová, alt. 1480 m, leg. J. Váňa, 1.09.1972 (herb. Váňa).

 

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