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Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014

Penaeid prawn resources along the east coast of India during 1991-2011 Maheswarudu, G., Sudhakara Rao G., Rajamani, M., Thangaraj Subramanian, V., Manmadhan Nair, K.R., Saleela, K.N., Dhanwanthari, E., Miriam Paul and Unnithan, A.K. Central Marine Fisheries Research Instiute, Kochi

The east coast of India with a coast line of 2,688 km and continental shelf area of about 0.56 million km2, provides a good habitat for the penaeid prawns in the adult phase. There are five major perennial rivers, number of creeks and low-lying areas offering nursery grounds for post larvae and juveniles that migrate to the sea after completion of nursery phase to get recruited to the fishery. Though as many as 23 species of penaeids are recorded along the east coast only about 19 species are supporting the regular fishery. Craft&gear Trawl nets are the major gearwhich is exploiting more than 90% of penaeid prawn along the east coast. Trawlers are of different types depending on the size of the boat, engine capacity and size of the gear. These are Pablo (9.14 m), Royya (9.75-10 m), Sorrah (11.4 m) and Sona (13.1 m) boats.The major fishing harbours/ landing centres, which are bases for operation of trawlers are Diamond harbor, Digha, Paradeep, Visakhapatnam, Kakinada, Chennai, Mandapam and Tuticorin. Besides these trawlers,Thalluvalai along the Tamil Nadu coast and stake nets along the Andhra Pradesh coast are also operated for exploitation of juvenile prawns in shallow coastal waters, estuaries, creeks and backwaters.

2,19,054 t in 2011. During 1991-2000 catch has fluctuated with 0.054 compound annual growth rate whereas during 2001-2010 sharp increase in catch was recorded with 0.156 compound annual growth rate. Overall increasing trend was observed during 21 years period (Fig.1). Average instead of mean state wise contribution of penaeid prawns shows Tamil Nadu contributed the highest (32%) followed by Andhra Pradesh (26%), West Bengal (22%),Odisha (19%), and Puducherry (1%). The highest catch was recorded from West Bengal and Odisha in 2011, from Andhra Pradesh in 2010 and from Tamil Nadu and Puducherry in 2009. The lowest catch was registered from West Bengal, Odisha, Andhra Pradesh in 1991, from Tamil Nadu in 2001 whereas from Puducherry it was in 1997 (Table1). West Bengal has the highest compound annual growth rate (0.211) followed by Odisha (0.192), Andhra Pradesh ( 0.056) and Tamil Nadu (0.033) during 21 years period, whereas Puducherry showed negative compound annual growth rate (-0.006). All states

Penaeid prawn landings Penaeid prawns, on average, contributed 9.8% of annual total marine fish landings along the east coast and theircontribution ranged from 5.5% to 13.8% during 1991 – 2011. On an average, east coast, contributed 29% of total penaeid prawn catch of India and its share ranged from 17.4% to 41.2% during the 21 years period. Average annual catch was 86,969 t and it ranged from 33,131 t in 1991 to

Fig. 1. Trends in penaeid landings and its share in total penaeid landings along the east coast

0.192

0.134

0.334

0.211

0.133

0.185

0.017 0.081

0.086

0.033

18523 20286 19833 30176 28038 27528 27284 28348 23443 22004 16202 21266 34723 32395 27146 35775 32078 35450 37127 35358 36840 589823 28087

Tamil Nadu

0.077

0.056

10759 10797 16200 15513 13863 15138 14193 19011 24967 22657 16221 16391 28382 26607 22158 30350 33214 30656 34001 36922 33877 471877 22470

Andhra Pradesh

CAGR: Compound annual growth rate

1972 2738 2986 2520 5350 3557 2966 2276 4323 6911 4105 4947 8826 12587 17293 13094 23077 29266 54453 73225 79111 355583 16933

1223 2677 2754 1247 3352 3799 3030 3123 2704 4272 8780 9434 36104 23905 29646 28135 28158 37703 49191 47952 68645 395834 18849

1991 1992 1993 1994 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 2005 2006 2007 2008 2009 2010 2011 Total Mean CAGR for 1991-11 CAGR for 1991-2000 CAGR for 2001-2010

Odisha

West Bengal

Year

0.197

-0.064

-0.006

654 400 146 785 458 361 104 702 368 339 154 653 221 704 248 1619 500 1441 1861 932 581 13231 630

0.156

0.054

0.094

33131 36898 41919 50241 51061 50383 47577 53460 55805 56183 45462 52691 108256 96198 96491 108973 117027 134516 176633 194389 219054 1826348 86969

Puducherry Total (east coast)

0.103

0.007

0.050

190210 189840 173204 224623 178874 187791 208540 216343 174071 204278 176448 203801 432571 362214 352776 426416 401688 488669 490722 471291 532851 6287221 299391

All India penaeid prawn landings

Table 1. Statewisepenaeid prawn landings (t) along the east coast during 1991-2011

0.048

0.047

0.042

17.4 19.4 24.2 22.4 28.5 26.8 22.8 24.7 32.1 27.5 25.8 25.9 25.0 26.6 27.4 25.6 29.1 27.5 36.0 41.2 41.1 29.0 29

Contribution of east coastto all India penaeid prawn landings (%)

0.039

0.017

0.026

2251255 2310052 1976143 2359525 2258832 2380842 2692409 2635670 2401706 2652928 2292703 2589645 2587095 2538105 2295490 2710988 2888461 3207205 3205453 3346687 3820207 55401401 2638162

Total marine fish landings

0.062

-0.009

0.024

8.4 8.2 8.8 9.5 7.9 7.9 7.7 8.2 7.2 7.7 7.7 7.9 16.7 14.3 15.4 15.7 13.9 15.2 15.3 14.1 13.9 11.3 11.3

Contribution of penaeid prawn landings to total marine fishlandings (%)

0.075

0.023

0.048

600917 660693 669937 688127 700222 747442 807752 758894 737464 751484 684856 809999 824638 885609 749385 866749 1033225 1146745 1450125 1411678 1599885 18585826 885039.3

Total marine fish landings of east coast

0.076

0.031

0.044

5.5 5.6 6.3 7.3 7.3 6.7 5.9 7.0 7.6 7.5 6.6 6.5 13.1 10.9 12.9 12.6 11.3 11.7 12.2 13.8 13.7 9.8 9.8

Cntribution of penaeid landings to total marine fish landings of the east coast

Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014 9

Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014

10

including Puducherry had higher compound annual growth rates during 2001-2010 than those during 1991-2000. Species Composition Species composition of penaeid prawn catches along the east coast was computed using the species composition data collected at Paradeep,

Visakhapatnam, Kakinada, Chennai, Mandapam and Tuticorin fishing harbours/ landing centres. Among the 23 species recorded 19 species had supported theregular fishery (Table 2). Metapenaeus dobsoni dominated the catch by contributing 21.4% followed by M. monoceros (3.2%), Metapenaeopsis spp (9.4%), Penaeus semisulcatus (9.2), Fenneropenaeus indicus (7.5%), Parapenaeopsis maxillipedo (4.3%),

Table 2. Common name and local names in four regional languages for penaeid prawns along the east coast S.No. Species

Common Name

Local name Oriya

Telugu

Tamil

1

Metapenaeus dobsoni

Kadalshrimp

Garangchingri

Khopra, Ranichingudi

Chinkiroyya

Chemakkaraeral

2

Metapenaeus monoceros

Speckled shrimp

Kara chingri

Khopra, Ranichingudi

Chakuroyya/ Kalandhan

Valuchaeral

3

Penaeus semisulcatus

Green tiger prawn

Bagda

Bagada, Katlareyya

Nooneroyya

Varieral

4

Peneaus indicus

Indian white shrimp

Chapra

Chapda, Tellareyya

Tellaroyya/ Narran

Vellaeral/ Vellaieral

5

Metapenaeus affinis

Jingashrimp

Chamneychingri

Khopra, Kalireyya

Gullaroyya/ Keliroyya

Chayavaluchaeral

6

Penaeus monodon

Giant tiger shrimp

Keleghari, Bagda

Bagada, Katlareyya

Katlaroyya

Karuvandueral/ Kathambaeral

7

Penaeus merguiensis

Banana shrimp

Chapda, Pettireyya

Kalliroyya

Vellaeral/ Vellaieral

8

Parapenaeopsis stylifera

Kiddi shrimp

Matka

Koddi, gullareyya

Gullaroyya / Karrkadi

Vandueral

9

Parapenaeopsis hardwickii

Spear shrimp

Lalchingri

Khodi, gullareyya

Gullaroyya/ Karrkadi

Vandueral

10

Metapenaeus lysianassa

Bird shrimp

—

—

—

Vellaiveluchaeral

11

Solenocera spp.

Coastal mudshrimp

—

Nallichingudi, Errareyya

Kukkaroyya

Kalleral

12

Metapenaeopsis spp.

Fiddler shrimp/ Velvet shrimp

—

—

Gullaroyya

Pottueral

13

Parapenaeopsis maxillipedo

Torpedo shrimp

—

—

Gullaroyya

Karikkada/ Vandueral

14

Parapenaeopsis uncta

Uncta shrimp

—

—

Gullaroyya

Vandueral

15

Trachypenaeus spp.

Rough shrimp

—

—

Garukugullaroyya Vandueral

16

Metapenaeus moyebi

Moyebi shrimp

—

—

—

—

17

Parapenaeus longipes

Flamingo shrimp

—

—

—

Thattaieral

18

Metapenaeus brevicornis

Yellow shrimp

Chamneychingri

Khopra, Kali reyya

Pasupuroyya / Puvvalin

Manjavalucheral

19

Penaeus japonicus

Kuruma shrimp

Kaonra, Pamra

Bagada, Katlareyya

Kalliroyya

Kathampaeral

20

Other penaeids

—

—

—

—

—

Bengali

1996 11281 5361 5080 3654 1484 811 174 2210 1962 796 2358 5104 2079 570 2401 2014 36 681 47 2280 50383 1995 9521 6636 2503 4116 2297 734 184 2288 2185 1271 2678 5494 2853 592 2770 1449 794 661 55 1980 51061 1994 11423 5611 3476 4402 1171 1389 177 2379 819 726 1650 4144 3730 524 2596 1391 674 1111 92 2757 50241 1993 10200 4818 4332 2928 1558 592 133 1321 1444 372 1738 3644 1551 343 1538 689 615 720 77 3306 41919 1992 7592 3540 4851 3130 1119 426 298 671 509 1054 545 4999 1363 217 1574 284 24 0 0 4702 36898 1991 6581 3384 4524 2926 1000 404 270 484 300 637 452 4634 1264 201 1459 264 22 0 0 4327 33131 Species Metapenaeus dobsoni Metapenaeus monoceros Peneausse misulcatus Peneaus indicus Metapenaeus affinis Peneaus monodon Peneaus merguiensis Parapenaeopsis stylifera Parapenaeopsis hardwickii Metapenaeus lysianassa Solenocera spp. Metapenaeopsis spp. Parapenaeopsis maxillipedo Parapenaeopsi suncta Trachypenaeus spp. Metapenaeus moyebi Parapenaeus longipes Metapenaeus brevicornis Penaeus japonicus Other penaeids Total 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20

Table 3. Species composition of penaeid landings by weight(t) along the east coast

1997 9044 5647 5520 3412 1275 616 173 2418 2122 762 1705 4732 1822 500 1830 2409 386 669 51 2485 47577

1998 13867 8744 4613 4361 1152 667 156 2173 2119 528 1982 4202 2551 489 1932 677 395 1167 37 1650 53460

1999 11986 10111 4347 3262 1498 791 195 2398 2557 821 2995 3884 1829 494 1741 2008 400 827 42 3619 55805

2000 10670 8854 4578 3485 2048 966 308 2525 3483 1175 3278 4015 1627 459 1949 921 1123 1571 64 2961 56061

2001 8035 5971 3644 2769 2407 791 322 2512 3801 1269 2938 3426 1227 425 1287 518 693 978 44 2404 45462

2002 9064 6619 4838 3519 2638 961 357 2754 4224 1408 3241 4165 1633 508 1697 665 758 989 44 2609 52690

Average 10217 6271 4382 3568 1460 739 207 1887 1750 814 1938 4485 2067 439 1979 1211 447 741 47 3007 47654

% 21.4 13.2 9.2 7.5 3.1 1.6 0.4 4.0 3.7 1.7 4.1 9.4 4.3 0.9 4.2 2.5 0.9 1.6 0.1 6.3 100.0

Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014

11

Trachypenaeus spp (4.2%), Solenocera spp (4.1%), Parapenaeopsiss tylifera (4%), P.hardwickii (3.7%), M. affinis (3.1%), M. moyebi (2.5%), M. lysiansa (1.7%), P. monodon (1.6%), M. brevicornis (1.6%) and Parapenaeus longipes. M. brevicornis has emerged as a regular species from 1993 onwards. Contribution of P. stylifera, P. hardwickii, Solenocera spp, P. longipes and M. moyebi had significantly increased from 1993 onwards. Threefold increase in the catch of P. monodon was observed in 1994 and thereafter its contribution was more or less stable. Contribution of F. indicus, M. dobsoni, M. monoceros and P. maxillipedo were gradually increased up to 1994 and since then their status was maintained. West Bengal and Odisha: About 11 species supported the fishery of which P. hardwickii (24.1%) dominated followed by M. dobsoni (18.5%), P. stylifera (11.7%), M. lysianassa (10.4%), Solenocera spp (7.9%) and M. monoceros (6.6%). Commercial species like P. monodon, F. indicus and P. merguiensis contributed only in low quantities (Table4). Andhra Pradesh: About 18 species supported the fishery. M. dobsoni (19.1%) dominated followed by M. monoceros (25.2%), Solenocera spp. (7.1%), M. brevicornis (4.8%) P. stylifera (4.1%), F. indicus (3.8%), M. affinis (3.3%), and M. spp (3.0%). Other highly commercial species like P. monodon (1.8%), P. semisulcatus (1.3%) and P. merguiensis (0.6%) were contributed in low volumes only. Tamil Nadu and Puducherry: About 16 species contributed to the fishery. P. semisulcatus contributed high (17.3%) followed by others (16.2%), M. dobsoni(15.8%), P. indicus(11.8%), P. maxillipedo (8.1%), Trachypenaeus spp (7.7%), M. monoceros (6.9%), and M. moyebi(4.5%). The commercial species like P. monodon (1.7%) and P. merguiensis (0.04%) were contributed in low quantities. This is the first report on species composition of penaeid prawn landings along the east coast of India from different maritime states such as West Bengal, Odisha, Andhra Pradesh, Tamil Nadu and Puducherry. Species composition, size range and modal groups of both sexes of commercial species at different fishing harbours/landing centres of present report can be utilized for future comparative studies.

1991 1151 114 0 73 128 11 33 242 300 597 102 0 0 0 0 0 0 0 0 444 3195

1992 1951 193 0 124 218 18 55 410 509 1011 173 0 0 0 0 0 0 0 0 752 5415

1993 1377 293 0 11 695 38 29 554 929 234 289 0 0 0 0 0 0 0 0 1291 5740

1994 849 113 0 5 551 22 25 402 486 443 160 0 0 0 0 0 0 0 0 710 3767

1995 1201 305 0 17 1349 35 78 1166 1906 1123 1096 0 0 0 0 0 0 0 0 426 8702

1996 1386 377 0 15 819 30 58 966 1638 722 819 0 0 0 0 0 0 0 0 524 7356

1997 1105 329 0 17 824 50 101 839 1590 625 343 0 0 0 0 0 0 0 0 173 5996

1998 1057 416 0 22 553 51 114 631 1630 415 376 0 0 0 0 0 0 0 0 134 5399

Species Metapenaeus dobsoni Metapenaeus monoceros Penaeus semisulcatus Penaeus indicus Metapenaeus affinis Penaeus monodon Penaeus merguiensis Parapenaeopsis stylifera Parapenaeopsis hardwickii Metapenaeus lysianassa Solenocera spp. Metapenaeopsis spp. Parapenaeopsis maxillipedo Parapenaeopsis uncta Trachypenaeus spp. Metapenaeus moyebi Parapenaeus longipes Metapenaeus brevicornis Penaeus japonicus Other penaeids Total

1991 2871 2414 397 952 510 216 184 0 0 0 68 0 0 0 0 0 0 0 0 3147 10759

1992 2881 2422 399 956 511 217 185 0 0 0 68 0 0 0 0 0 0 0 0 3158 10797

1993 5669 3195 290 705 674 285 105 612 515 120 1418 377 0 0 16 0 187 720 77 1234 16200

1994 4264 3058 340 458 571 1010 152 1006 333 225 1228 396 0 0 0 0 34 1111 92 1237 15513

1995 3573 4086 170 686 496 328 106 664 279 84 1404 105 0 0 143 0 148 661 55 876 13863

1996 5904 3311 78 596 455 257 114 616 324 74 1121 509 0 0 0 0 36 681 47 1015 15138

1997 4120 3607 160 455 299 184 70 1008 531 138 982 223 0 0 0 0 6 669 51 1691 14193

1998 6758 6173 113 485 421 74 41 772 488 113 1052 322 0 0 0 0 41 1167 37 954 19011

Table 5. Species wise landings (t) of penaeid prawns along the Andhra Pradesh coast

Species Metapenaeus dobsoni Metapenaeus monoceros Penaeus semisulcatus Penaeus indicus Metapenaeus affinis Penaeus monodon Penaeus merguiensis Parapenaeopsis stylifera Parapenaeopsis hardwickii Metapenaeus lysianassa Solenocera spp. Metapenaeopsis spp. Parapenaeopsis maxillipedo Parapenaeopsis uncta Trachypenaeus spp. Metapenaeus moyebi Parapenaeus longipes Metapenaeus brevicornis Penaeus japonicus Other penaeids Total

Table 4. Species wise landings (t) of penaeid prawns along West Bengal and Odisha coasts

1999 7297 8104 132 497 678 302 42 954 445 106 2064 549 0 0 0 0 128 827 42 2800 24967

1999 1178 533 0 28 685 72 151 852 2112 715 507 0 0 0 0 0 0 0 0 195 7027

2000 5675 6016 204 581 410 242 48 825 336 143 1750 827 26 182 127 0 873 1571 64 2672 22573

2000 1617 961 0 82 1373 151 261 1323 3147 1032 972 0 0 0 0 0 0 0 0 265 11183

2001 3903 3443 135 517 708 184 21 821 175 80 1352 1290 36 188 86 0 518 978 44 1742 16221

2001 1863 1107 0 94 1582 174 301 1524 3626 1189 1119 0 0 0 0 133 0 0 0 171 12885

2002 3944 3479 136 523 716 186 21 830 177 81 1366 1304 36 190 86 0 524 989 44 1760 16391

2002 2079 1236 0 105 1766 195 335 1701 4047 1327 1249 0 0 0 0 149 0 0 0 191 14381

Total 56859 49310 2553 7413 6449 3485 1089 8107 3603 1162 13871 5900 99 560 458 0 2496 9373 553 22284 195626

Total 16814 5978 0 594 10543 848 1542 10610 21921 9432 7205 0 0 0 0 282 0 0 0 5277 91046

Mean 4738 4109 213 618 537 290 91 676 300 97 1156 492 8 47 38 0 208 781 46 1857 16302

Mean 1401 498 0 50 879 71 128 884 1827 786 600 0 0 0 0 23 0 0 0 440 7587

% 29.1 25.2 1.3 3.8 3.3 1.8 0.6 4.1 1.8 0.6 7.1 3.0 0.1 0.3 0.2 0.0 1.3 4.8 0.3 11.4 100.0

% 18.5 6.6 0.0 0.7 11.6 0.9 1.7 11.7 24.1 10.4 7.9 0.0 0.0 0.0 0.0 0.3 0.0 0.0 0.0 5.8 100.0

12 Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014

1991 2559 857 4127 1900 361 176 54 242 0 40 282 4634 1264 201 1459 264 22 0 0 736 19177

1992 2760 924 4452 2050 390 190 58 261 0 43 304 4999 1363 217 1574 284 24 0 0 792 20686

1993 3154 1330 4041 2211 190 269 0 155 0 18 31 3268 1551 343 1522 689 428 0 0 781 19979

1994 6309 2439 3137 3938 50 357 0 972 0 58 262 3748 3730 524 2596 1391 640 0 0 810 30961

1995 4748 2245 2332 3413 452 371 0 458 0 65 178 5389 2853 592 2627 1449 646 0 0 678 28496

1996 3991 1673 5002 3043 209 523 2 628 0 0 418 4595 2079 570 2401 2014 0 0 0 740 27889

1997 3819 1711 5359 2940 152 381 2 571 0 0 380 4509 1822 500 1830 2409 380 0 0 621 27388

1998 6052 2154 4500 3854 178 542 0 769 0 0 554 3881 2551 489 1932 677 354 0 0 561 29050

1999 3511 1474 4215 2736 135 417 1 593 0 0 424 3335 1829 494 1741 2008 272 0 0 625 23811

2000 3378 1877 4375 2822 265 573 0 377 0 0 555 3188 1601 277 1823 921 250 0 0 24 22305

2001 2269 1421 3509 2157 117 433 0 166 0 0 467 2136 1191 237 1202 385 174 0 0 491 16356

2002 3041 1904 4702 2891 157 580 0 223 0 0 626 2862 1597 318 1611 516 234 0 0 658 21919

Total 45593 20008 49752 33957 2655 4813 117 5415 0 224 4483 46543 23430 4760 22318 13008 3425 0 0 7516 288017

Paradeep P. monodon Male Female P. merguiensis Male Female M. affinis Male Female M. monoceros Male Female M.dobsoni Male Female

Landing centre & species

210-220 241-250

146-150 161-165

116-120 131-135

121-125 131-135

0 0

116-170 116-205

71-105 71-185

76-140 76-195

0 0

1992 Mode

91-250 91-300

Size range

Size range

1994 Mode

0 0

81-145 76-185

66-155 66-180

0 0

0 0

111-115 76-150 136-140 71-185

106-110 71-170 126-130 71-185

0 0

0 0

111-115 0 141-145 0

111-115 0 116-120 0

116-180 141-145 111-170 136-140 0 121-195 151-155 111-205 151-155 0

0 0

0 0

0 0

0 0

0 0

201-210 101-250 211-220 241-250 201-290 241-250

1997 Size Mode range

46-100 51-115

76-145 71-190

66-160 71-185

76-80 91-95

46-100 41-110

111-115 71-145 136-140 71-180

111-115 76-155 126-130 71-185

71-75 81-85

111-115 131-135

111-115 126-130

111-170 146-150 111-180 146-150 111-195 161-165 111-205 151-155

91-250 91-290

1995 1996 Size Mode Size Mode range range

201-210 91-250 211-220 0 241-250 101-300 171-180 0

1993 Mode

91-240 91-300

Size range

Size range

46-100 46-115

71-140 71-170

71-155 71-185

66-70 81-85

41-100 41-115

111-115 76-150 131-135 71-185

116-120 71-155 131-135 71-180

111-170 146-150 111-180 111-200 161-165 111-210

2000 Size Mode range

Size range

Mean 3799 1667 4146 2830 221 401 10 451 0 19 374 3879 1952 397 1860 1084 285 0 0 626 24001

81-85 76-80

41-100 71-75 51-110 81-85

0 0

116-120 81-150 111-115 0 131-135 71-185 131-135 0

111-115 71-160 106-110 0 126-130 71-185 126-130 0

146-150 111-170 146-150 0 161-165 111-215 136-140 0

201-210 101-240 201-210 0 211-220 101-300 231-240 0

1999 Mode

201-210 101-260 241-250 91-290

1998 Mode

91-240 91-290

Size range

Table 7. Size range (mm) and mode (mm) of commercial species of penaeid prawns at different landing centres along the east coast.

Species Metapenaeus dobsoni Metapenaeus monoceros Penaeus semisulcatus Penaeus indicus Metapenaeus affinis Penaeus monodon Penaeus merguiensis Parapenaeopsis stylifera Parapenaeopsis hardwickii Metapenaeus lysianassa Solenocera spp. Metapenaeopsis spp. Parapenaeopsis maxillipedo Parapenaeopsi suncta Trachypenaeus spp. Metapenaeus moyebi Parapenaeus longipes Metapenaeus brevicornis Penaeus japonicus Other penaeids Total

Table 6. Species wise landings (t) of penaeid prawns along the Tamil Nadu and Puducherry coasts

0 0

0 0

0 0

0 0

0 0

2001 Mode

% 15.8 6.9 17.3 11.8 0.9 1.7 0.04 1.9 0.0 0.1 1.6 16.2 8.1 1.7 7.7 4.5 1.2 0.0 0.0 2.6 100.0

Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014 13

Male Female P.indicus Male Female P.monodon Male Female M.dobsoni Male Female Chennai M.dobsoni Male Female P.maxillipedo Male Female Mandapam P.semisulcatus Male Female Tuticorin P.semisulcatus Male Female

Visakhapatnam M.monoceros Male Female P. indicus Male Female P.semisulcatus Male Female Kakinada M.monoceros

126-130 121-125

181-190 181-190

66-70 76-80

71-75 81-85

81-85 81-85

126-130 131-135

0 0

76-185 51-215

91-260 91-300

41-105 41-115

46-105 46-110

46-105 46-105

76-200 71-215

101-172 91-226

0 0

0 0

81-85 76-80

161-165 181-185

111-195 116-210

46-165 41-175

111-115 146-150

76-160 81-200

126-130 86-170 146-150 96-205

126-130 86-170 0 161-165 96-200 0

95-188 106-205

76-165 76-205

41-105 41-115

46-150 46-110

0 0

0 0

0 0

0 0

0 0

0 0

116-120 126-130

76-80 81-85

71-75 86-90

0 0

0 0

0 0

0 0

0 0

86-175 86-235

86-160 86-200

46-105 41-115

46-105 41-105

41-100 41-110

86-230 101-270

86-180 81-200

41-145 36-180

126-130 161-165

116-120 116-120

71-75 86-90

81-85 86-90

66-70 76-80

156-160 161-165

116-120 126-130

86-90 86-90

0 0

0 0

0 0

0 0

0 0

0 0

0 0

0 0

0 0

126-130 93-168 141-145 93-203

121-125 146-150

0 0

0 0

0 0

101-230 101-280

81-180 51-200

51-160 46-190

0 0

96-160 96-225

121-125 161-165

106-110 126-130

0 0

0 0

0 0

171-180 191-200

121-130 121-130

66-70 76-80

0 0

0 0

91-150 96-200

0 0

0 0

41-100 41-110

101-250 81-270

81-180 81-270

46-170 46-185

0 0

116-120 126-130

0 0

0 0

76-80 76-80

161-170 161-170

131-140 131-140

91-95 91-95

133-223 171-175 138-238 201-205

121-205 161-165 103-198 166-170 121-215 166-170 123-208 186-190

86-165 86-200

91-155 111-115 81-150 96-200 141-145 86-200

0 0

0 0

0 0

0 0

0 0

0 0

126-215 181-185 0 131-250 231-235 0

101-200 146-150 121-195 166-170 121-195 0 81-210 166-170 131-230 186-190 131-230 0

86-185 86-210

121-125 98-178 141-145 83-203

0 0

86-150 86-190

0 0

46-110 41-115

31-100 41-110

91-220 91-240

81-170 71-190

46-130 41-155

0 0

106-110 116-120

0 0

66-70 66-70

66-70 66-70

181-190 181-190

131-140 131-140

81-85 81-85

0 0

91-160 91-205

0 0

0 0

31-95 31-115

81-240 101-300

71-180 71-200

51-145 51-165

113-228 176-180 0 93-238 211-215 0

113-193 156-160 113-193 128-238 166-170 113-238

93-168 93-208

0 0

116-120 136-140

0 0

0 0

66-70 81-85

151-160 151-160

121-130 121-130

86-90 86-90

0 0

121-130 121-130

81-85 86-90

0 0

96-160 96-205

0 0

0 0

41-90 41-110

0 0

111-115 126-130

0 0

0 0

66-70 81-85

101-240 151-160 111-280 161-170

81-170 81-180

51-130 51-160

96-190 101-235

96-150 101-200

0 0

46-110 41-115

41-90 36-115

91-230 91-290

81-180 81-190

46-135 46-170

116-120 151-155

121-125 126-130

0 0

61-65 61-65

66-70 66-70

131-140 151-160

121-130 121-130

81-85 91-95

111-200 166-170 101-240 176-180 106-255 191-195 106-260 206-210

161-165 126-190 156-160 121-195 151-155 186-190 121-205 151-155 106-205 166-170

136-140 81-180 136-140 71-175 136-140 161-165 91-215 161-165 71-205 131-135

14 Mar. Fish. Infor. Serv., T & E Ser., No. 219, 2014