Acta Zoologica Academiae Scientiarum Hungaricae 47 (4), pp. 349–361, 2001

A NEW GENUS AND THREE NEW SPECIES OF HYBOTIDAE WITH NEW RECORDS OF THE HUNGARIAN EMPIDOIDEA (DIPTERA) PAPP, L. and M. FÖLDVÁRI Department of Zoology, Hungarian Natural History Museum H-1088 Budapest, Baross u. 13, Hungary E-mail: [email protected], [email protected] Twenty-two species are recorded as new for the Hungarian fauna, among them three species (Tachypeza yinyang sp. n., Megagrapha europaea sp. n., Chvalaea sopianae sp. n.) are new to science. The genus Megagrapha is reported from the Palaearctic region for the first time. A new genus of Ocydromiini, Chvalaea gen. n. (type species Leptopeza rugosiventris STROBL, 1910) is described with two species from Hungary. Key words: Chvalaea, Megagrapha, Hybotidae, Microphoridae, Empididae, faunistic survey, new records, Hungary

INTRODUCTION In the frame of the project “Large blank spots in the Diptera fauna of Hungary” we are to collect and to publish species representing dipterous families formerly not recorded from Hungary. Furthermore, species representing not recorded genera are also targets of our activity. Four years of the project (1999–2002) comprise a reasonable period of time, for which a fair support is provided for true faunistical studies. The results are included also in the “Checklist of the Diptera of Hungary” (PAPP et al. 2001). In the first two years of the project these collectings resulted in capturing specimens of numerous genera and species new for the fauna of Hungary. We published a paper with some new records on Empidoidea last year (PAPP & FÖLDVÁRI 2000), further results are reported in this paper. All the specimens below are deposited in the Diptera collection of the Department of Zoology, Hungarian Natural History Museum, Budapest (below: HNHM). Some abbreviations used below and also on collection labels: NP: Nemzeti Park [National Park], TK: Tájvédelmi Körzet [Landscape Protected Area], TT: Természetvédelmi Terület [Nature Reserve], hg., hegys.: hegység [mountains]. The following Hungarian words are on numerous labels: “csapda”: trap, “erdő”: forest, “rét”: meadow, “patak fölött”: over the brook, “patak fölött és mellett”: over and alongside the brook; völgy, v: valley; p, patak: brook. The months are given as on the collection labels, i.e., May: május, 05., June: június, 06., July: július, 07., October: okt., 10., etc.; since labels are written in Hungarian, months come first. Acta zool. hung. 47, 2001 Hungarian Natural History Museum, Budapest

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HYBOTIDAE Megagrapha europaea sp. n. (Figs 1–5) Holotype male: Kelet-Mecsek TK, Komló: Zobákpuszta, Hidasi-völgy – patak fölött, mellett, 2000. június 13., leg. PAPP L. Paratypes: 1 male, 2 females: same as for holotype; 1 female: Zempléni TK: Regéc, Ördög-v., patak fölött és mellett, 2000. július 5., leg. PAPP L.; 1 male: Kelet-Mecsek TK: Óbánya, Óbányai-p. fölött és mellett, 2000. június 14., leg. PAPP L. (wings prepared on a slide, body with genitalia in a plastic microvial with glycerine, head and left fore leg glued on a small card under the microvial, right fore tibia and tarsi lost). Measurements in mm: body length 1.54 (holotype), 1.50–1.55, 1.71–1.83 (paratype males, females), wing length 1.96 (holotype), 1.83–1.95, 2.05–2.25 (paratypes), wing breadth ca. 0.75 (holotype), 0.82, 0.79–0.96 (paratypes, not measurable on one of the males). Male head and abdomen black, thorax yellow but pronotal region with proepisternal depression, and metanotum, scutellum with a prescutellar mesonotal area, also black. Female body all black. Head actually without characteristic setae: ocellars hairlike, vertex and occiput with a number of thin white hairs. Eyes meet in the facial area, eyes densely and comparatively long pubescent. Frons (praefrons) shiny, very narrow anteriorly. Scape small, pedicel somewhat larger, first flagellomere ca. 1.5 times longer than broad (0.086 mm vs 0.055 mm), i.e. not as in M. platytarsis, see fig. 4 of CHILLCOTT and TESKEY (1983). Arista (Fig. 2) 0.26 mm long, somewhat thickened with dense, long cilia. Thorax without characteristic setae, or rather, they are not longer and thicker than the enlarged microchaetae. In any case, 1 longer prescutellar dorsocentral pair and a row of 4 pairs of notopleurals (all pale yellow) are discernible. Scutellum with 4 pairs of marginal sc, which emerge on small tubercles. Legs clear yellow, incl. tarsi. Legs without armature, male mid tarsi normal (Fig. 3), mid basitarsus 1.5 times longer than second tarsomere (the latter not broadened). Wings clear with long microtrichia (Fig. 1), veins yellow. Costal cilia up to 0.085 mm. Subcostal vein weak, discontinued at about its distal 3/5 towards costa. Vein R1 short, costal sections 2:3:4 = 85:69:30. Vein R4+5 slightly upcurved in its basal part, but apically downcurved, consequently, 4th costal section much shorter than 1/2 length of the 2nd section. R-M and dM-Cu cross-veins meet, their length ratio is 3 to 5 (R-M and their meeting point discoloured). Distal section of the cubital vein 0.60 mm, i.e. considerably longer than proximal section. Anal vein thin but discernible and continued along a wide arc, rather close to the wing margin, well distally to the dM-Cu cross-vein. Alula small with long bristles. Knob of halteres black, stalk yellowish. Abdominal terga and sterna weakly sclerotized. Male genitalia with fused and asymmetrical cerci, with asymmetrically placed long apical setae (Figs 4–5). Right lamella narrow, digitiform in profile, left lamella very large, continued ventrally, with a narrow recurving process subbasally (Fig. 4). Female cerci medium-long with short hairs only.

Similar to M. platytarsis CHILLCOTT, 1983 but smaller (wing length 1.83 to 2.25 mm, vs. 2.4–2.6 mm that of M. platytarsis), its male mid second tarsomere is much shorter than first one (basitarsus), and knob of halteres is dark, not yellow as Acta zool. hung. 47, 2001

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in M. platytarsis. The male first flagellomere is strongly and also costal ratios are slightly different from those of M. platytarsis. Etymology. This is the first known species of the genus Megagrapha Melander, 1928 from Europe (and also from the Palaearctic region).

Figs 1–5. Megagrapha europaea sp. n., paratype male. 1: wing; 2: arista; 3: male mid leg, anterior view; 4: genitalia, dorsolateral view; 5: genitalia, dorsal view. Scales: 1.0 mm for Fig. 1; 0.2 mm for Fig. 2; 0.3 mm for Figs 4–5 Acta zool. hung. 47, 2001

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Tachypeza yinyang sp. n. (Figs 6–10) Holotype male: K-Mecsek TK: Óbánya, Óbányai-völgy, patak fölött, 1999. május 30., leg. PAPP LÁSZLÓ. Paratypes: 1 male (abdomen with genitalia in a plastic microvial with glycerine): Melegmányi-völgy TT: Pécs, Nagy-mély-v., patak fölött, 1999. május 27., leg. PAPP LÁSZLÓ; 1 male (teneral but intact specimen): Kőszegi TK: Kőszeg, Hármas-p. fölött és mellett, 2000. június 28., leg. PAPP L. Measurements in mm: body length 3.33 (holotype), 2.50, 3.30 (paratypes), wing length 3.58 (holotype), 3.21, 3.70 (paratypes), wing breadth 1.18 (holotype), 1.00, 1.17 (paratypes). Body grey microtrichose but pleura darker with distinct reflection. Head higher than long, occiput convex, antennae inserted slightly dorsally to middle of head seen in profile. Antennae yellow, arista microtrichose, 0.80 mm long. Occiput with 2 pairs of thick short setae: 0.10 mm and 0.075 mm long; plus a wreath of medium-long whitish setae around the neck and posteriorly on gena (like in T. truncorum). Palpi with pale terminal bristles. Only 1 (posterior) pair of notopleurals and a pair of strong, widely separated scutellars; both pairs are very thick. Acrostichal and dorsocentral microchaetae are scarce and almost indiscernible. No dorsocentrals, in the place of posterior dc-s a pair of thin short hairs (only 0.02 mm long!). Scutellars (paratype) 0.16 mm long. Legs mostly yellow, mid and hind coxae blackish, apical tarsomeres darkened; tibial and femoral structures and armature are similar to those of T. truncorum. Fore coxae without black spines, yellow, anterior apical 3/5 with a black stripe, which widens apically. Fore tibia robust (Fig. 6), both laterally and medially (posteriorly and anteriorly) with a black band: anteriorly in apical half, posteriorly in apical 3/5. Anterior surface of fore femora (Fig. 6) medially with sharply margined black pattern, which resembles to the Chinese yin and yang; fore femur without long dark hairs subapically (as in T. truncorum). Mid femur (Fig. 7) black in proximal third, dilated basally with a slight emargination subbasally, this basal part with 5 or 6 spine-like setae ventrally. Mid tibia with a larger emargination subapically (Fig. 7). Hind femur thin, black basally, which turns to dirty yellow apically. Hind tibia black in its apical fifth. Wings light greyish with distinct brown hue along R1, R4+5 and Cu veins and also by anal vein stump (CuA2); brown colour widens into an obscure spot at fork of R2–5. Vein M strongly upcurving, conjointment with costa is before wing apex. Costa and apical half of vein R1 merge into a blackish brown flat structure. Male cerci (Fig. 9) strongly asymmetrical: right cercus simple, left cercus with a rather sharp apical projection medially and with another process ventro-laterally. Also the bristles are asymmetrically placed on cerci. Right paramere (Fig. 8) very large in three lobes, medial one with stronger setae. Left lamella (Fig. 10) rather large but more simple, laterally-subapically with a short but sharp incision. Female unknown.

Tachypeza yinyang sp. n. is a very easily recognisable species. Actually it does not fit into the key of CHVÁLA (1975): its thoracic pleura are with a distinct reflection (like those of T. fuscipennis) but its scutellar setae are widely separated. It has only one pair of (posterior) notopleurals and no dorsocentrals. The black patActa zool. hung. 47, 2001

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tern on the anterior surface of fore femur and tibia, the structure and colour of mid femur and tibia, as well as the single pair of notopleurals and the lack of dorsocentral setae make it conspicuous. Its male genitalia are also distinctive. Tachypeza fennica TUOMIKOSKI, 1932 – 1 male: Bakonyszentlászló, Vinye, Cuha-szurdok, 2000. július 26., leg. PAPP [L.], BAJZA [Zs.]. New to Hungary. Tachydromia carpathica CHVÁLA, 1966 – 1 male, 1 female: Kelet-Mecsek TK, Komló: Zobákpuszta, Hidasi-völgy, patak fölött és mellett, 2000. július 13., leg. PAPP L. 1 male: Zempléni TK: Regéc, Ördög-völgyi patak fölött és mellett, 2000. július 3., leg. PAPP L. New to Hungary. Tachydromia woodi (COLLIN, 1926) – 1 male: Lasztonya [Zala megye], 1967. VI. 27., leg. MÓCZÁR L. New to Hungary.

Figs 6–7. Tachypeza yinyang sp. n., paratype male. 6: fore femur and tibia, anterior view; 7: mid femur and tibia, anterior view. Scale: 0.5 mm Acta zool. hung. 47, 2001

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Dysaletria nigripennis CHVÁLA, 1975 – 1 male, 1 female: Budapest, Pestszentlőrinc, Péterhalmi-erdő, erdei tisztás [forest clearings], 2001. 04. 29–30., leg. PAPP L. The genus and species are new for the Hungarian fauna. This species was described from Slovakia (Plešany, Král’ovský Chlmec) not far from the northern border of Hungary. Trichina pallipes (ZETTERSTEDT, 1838) – 2 males, 1 female: Duna-Ipoly NP: Szokolya, Szén-p. fölött és mellett, 2000. május 13., leg. PAPP L. First record from Hungary. Trichinomyia flavipes (MEIGEN, 1830) – 1 female: Aranyosgadány, hullott körtéről [fallen pear], 1970. VII. 30., leg. PAPP L.; 1 female: Újhuta, Zoltán-forr. [spring], 1960. V. 8., leg. Z SEBESS; 2 females (without heads): Kőkapu, forrás és forrásláp [spring and peat-bog], 1960. V. 7., leg. Z.SEBESS. WÉBER (1983) published it from the Barcs Juniper Woodland Nature Reserve, but we did not find the voucher specimen (male) in any of the materials we had got from the Pécs University. Consequently, one may regard the above data as the first reliable record from Hungary. Oedalea apicalis LOEW, 1859 – 1 female: Duna-Ipoly NP: Szokolya, Szén-p. fölött és mellett, 2000. május 13., leg. PAPP L. First record from Hungary. Oedalea hybotina (FALLÉN, 1816) – 1 female: Gagyvendégi, akácos széle [edge of a blacklocust (Robinia) forest], 2000. 05. 16., leg. PAPP L. First record from Hungary.

Chvalaea gen. n. (Figs 11–16) Type species: Leptopeza rugosiventris STROBL, 1910 Gender: feminine. Eyes bare and not meeting above antennae (dichoptic), but meet along their whole length below eyes (no postfrons, “face”), dorsal ommatidia of the same size as ventral ones in females. Frons (praefrons) small and rather narrow. Eyes large, gena very narrow, occiput convex (but not strongly), covered by numerous medium-long setulae. No characteristic setae on head. First flagellomere more or less long and narrow with pin-like arista (Figs 13–14). Proboscis very short. Mesonotum extremely convex (bulging), pronotum comparatively very large (Fig. 11), which make their habitus peculiar: if abdomen placed horizontally, head is ventral to thorax*. Scutellum is reduced to a small, blunt transverse process dorsally to a large metanotum. Laterotergite bare. Legs with long slender setae, which are two or even three times as long as the femoral or tibial diameter at their emergence. Fore coxa short (without peculiar setae), fore, mid and hind coxae not far from each other. Legs are not raptorial as for the structure of femora and tibia, but tarsi tend to incurve with strong 5th segment and claws. Femora and tibiae long, slender. Swelling of the fore tibial gland well-discernible on medial basal surface. Wings clear, long and comparatively narrow. Wings without axillary angle, anal lobe or alula (Figs 11–12). Costa without erect setae, nor other veins setose. Subcosta incomplete, not reaching costa. Rs originating much distally to level of cross-vein H, R-M at about middle of wing. Vein R4+5 terminates at apex of wing. Basal radial and basal medial cells are very long, basal medial cell termi-

*

According to our friend, Dr BÉLA DARVAS (pers. comm.) Chvalaea specimens must fly with their abdomen lowered in order to have their ocelli dorsal, which is vital for perception of any movement over the animal, i.e. for its survival. If this is true, their legs are most anteral when flying.

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nates well distal to R-M. Cell dm very long and distally placed. Two veins emitted from cell dm (M1 lost), which do not reach wing margin. Vein CuA2 oblique and S-shape curved, posterior cubital cell only half as long as basal medial cell. Anal vein long, wavely curved, ends free somewhat distal to the middle of wing. Abdomen long, straight (stiff), subcylindrical, abdominal sclerites heavily sclerotized, punctate. Female cerci short and hidden in the 8th segment. Since male is unknown, after the capture of a male it will be necessary to supplement this description with the features of the male genitalia.

Figs 8–10. Tachypeza yinyang sp. n., paratype male. 8: right lamella, broadest view; 9: cerci, broadest view; 10: left lamella, broadest view. Scale: 0.2 mm Acta zool. hung. 47, 2001

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This is one of the genera, which were mentioned as “undescribed genus B” in SINCLAIR and CUMMING (2000). We do not think this new genus would be closely related to Oropezella COLLIN; only the long cell dm and shape of the basal part of wing are shared characters. Etymology. We name this new genus after Dr MILAN CHVÁLA (Charles University, Prague) for his unparalleled achievements in the taxonomy of the family Hybotidae and other families of Empidoidea.

Chvalaea rugosiventris (STROBL, 1910) (Figs 14, 16) Material studied: 1 female: Melegmányi[-völgy] TT: Pécs, Melegmányi-völgy, patak fötött és mellett, 2000. június 15., leg. PAPP. Body length (measured in the thorax-abdomen axis) 4.30 mm, wing length 4.00 mm, wing breadth 1.14 mm. Body colours as in Ch. sopianae. Body ratios even more extreme than in Ch. sopianae, since the abdomen 3.32 mm long, thorax 0.98 mm long. Head 0.59 mm long. First flagellomere only slightly longer than scape and pedicel combined; antenna 0.405 mm long, arista curved (i.e. not precisely measurable, but longer than 0.55 mm, i.e. actually probably somewhat longer than 0.60 mm (Fig. 14). Pronotum 0.17 mm, metanotum 0.31 mm if measured dorsally. Legs long, slender, e.g. hind femur 1.54 mm, with long slender bristle: longest bristle on hind tibia 0.22 mm, diameter of tibia there 0.12 mm, longest bristle on mid tibia 0.28 mm, diameter of tibia there 0.09 mm. Fore coxa dark, short, 0.35 mm, hind coxae yellow, middle coxae intermediate (but as short as hind ones). Hind femur with a badly defined dark subapical ring, hind tibia dark in its apical 3/5 part. Tarsi wholly spirally incurved with strong claws, tarsomeres darker. Wings clear, long and comparatively narrow (length/ width ratio 3.5!). Wing otherwise similar to that of Ch. sopianae. Abdomen extremely long, straight, abdominal sclerites heavily sclerotized, punctate. Longest (submarginal) abdominal hairs 0.40 mm long. Female 8th segment cylindrical (Fig. 16), 0.56 mm long, so longer than that of Ch. sopianae, but its dorsal caudal swelling indistinct (caudal edge rather upcurved). First record for Hungary.

Chvalaea sopianae sp. n. (Figs 11–13, 15) Holotype female: Melegmányi[-völgy] TT: Pécs, Melegmányi-völgy, patak fölött és mellett, 2000. június 15., leg. PAPP. Paratypes: 2 females: same as for holotype. Measurements in mm: body length (from the most anterior point of thorax to the end of abdomen) 4.38 (holotype), 4.38, 4.47 (paratypes), wing length 4.28, 4.26, 4.32, wing breadth 1.28, 1.28, 1.34. Acta zool. hung. 47, 2001

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Body peculiar: head ventral to the thorax-abdomen axis (Fig. 11), thorax dark brownish grey, slightly dusted, postpronotal callus and postalar callus diffuse reddish, abdomen shiny black, rugose (punctate). Body ratios somewhat less extreme than in Ch. rugosiventris, since the abdomen 3.32–3.36 mm long, thorax in the abdominal axis 1.10 mm long. Legs mostly yellow but knees dark. Head 0.51–0.59 mm long. Eyes dichoptic and bare, but meet along their whole length below eyes, dorsal ommatidia of the same size as ventral ones. Frons (praefrons) small and as narrow at ocelli as diamater of fore knee. Gena very narrow, occiput convex, covered by a number of light, medium-long setulae. No characteristic setae on head. Palpi extremely short, only 0.07 mm, also proboscis very short, 0.10 mm. Scape and pedicel only slightly longer than half length of first flagellomere, the whole antenna slightly shorter than arista (Fig. 13): antenna of the holotype 0.517 mm, arista 0.56

Fig. 11. Chvalaea sopianae sp. n., paratype female, habitus Acta zool. hung. 47, 2001

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mm, ratio 1.08 (one of the paratypes: 0.53 mm, 0.57 mm, broken off on the other one). First flagellomere definitely longer than that of Ch. rugosiventris. Mesonotum extremely convex (bulging), pronotum 0.18 mm, metanotum 0.33 mm if measured dorsally. Scutellum minute, laterotergite bare. Thoracic setae much reduced: only 1 weaker postpronotal (humeral), which is 0.10 mm on one paratype; 1 pair of stronger posterior notopleurals (0.22 mm long on one paratype), no more characteristic setae on thorax. Legs long, slender, e.g. hind femur 1.54 mm, with long slender bristle: longest bristle on hind tibia 0.26 mm, diameter of tibia there 0.12 mm, longest bristle on mid tibia 0.26 mm. Fore coxa dark, short, 0.38–0.40 mm, hind coxae yellow, middle coxae intermediate (but as short as hind ones).

Figs 12–16. 12: Chvalaea sopianae sp. n., female wing. 13–14: female antenna, medial view: 13: Ch. sopianae sp. n.; 14: Ch. rugosiventris (STROBL). 15–16: female 8th abdominal segment, lateral view: 15: Ch. sopianae sp. n.; 16: Ch. rugosiventris (STROBL). Scales: 1.0 mm for Fig. 12, 0.5 mm for Figs 13–16 Acta zool. hung. 47, 2001

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Swelling of the fore tibial gland rather large, tibia at widest 2 times as broad as subapically. Hind femur with a badly defined dark subapical ring, hind tibia mostly dark, at most basally lighter. Tarsi wholly spirally incurved with strong claws, yellowish, hind tarsomeres darker. Wings clear (Fig. 12), long and comparatively narrow (length/width ratio 3.27–3.47). Wing otherwise similar to that of Ch. rugosiventris. Halteres waxy yellow, 0.71–0.77 mm long, distal part of stalk very thin. Abdomen extremely long, straight, abdominal sclerites heavily sclerotized, punctate. Longest (submarginal) abdominal hairs 0.36 mm long. Eighth abdominal segment much shorter than that of Ch. rugosiventris: female 8th segment (Fig. 15), 0.91 mm long measured at middle height, dorsocaudally with a large globular swelling (Fig. 15) (caudal edge 0.64 mm high). Last visible sternite (most likely the 9th) with dense white hairs (Fig. 15). Female cerci must be short, hidden in the 8th segment. All the three individuals are slightly damaged: wings of the holotype are splitted; left wing of one of the paratypes was much wrinkled, now glued on a card below the specimen in a less wrinkled position; antennae of the other paratype are lost, right wing of this specimen is mounted on a slide.

Based on STROBL’s (1910) description of Leptopeza rugosiventris, SINCLAIR and CUMMING (2000) were able to point out that the species must belong to

an undescribed genus. However, the description itself: “Die Legeröhre bildet eine dicke, ebenfalls grobpunktierte Röhre von der Länge der halben Breite des letzten Ringes...” and “Die Fühler sind etwas länger als das 3. Fühlerglied”, was not enough to make distinction between the two species as L. rugosiventris was captured together with the females of another species. In any case, to adduce full proof, it was necessary to study the type of L. rugosiventris. Dr MILAN CHVÁLA was kind enough to study the type of L. rugosiventris in STROBL’s Collection for us (pers. comm.). The new species Ch. sopianae is easily distinguishable from Ch. rugosiventris by its antennae and by the shape of its 8th abdominal segment.

Etymology. The specific epithet “sopianae” (noun) is the old Roman name of the city Pécs, where this new species originates from.

MICROPHORIDAE Microphor crassipes MACQUART, 1827 – 2 males: [Duna-Ipoly NP] Diósjenő, Kemence-patak, 1995. VI. 20., leg. PAPP L., Ádám L. A species new for the Hungarian fauna (this is the third known species in our country).

EMPIDIDAE Hilara albiventris VON ROSER, 1840 – 1 male: Zempléni TK: Nagyhuta, Kemence-patak égeresei [forest with Alnus], 2000. júl. 6., leg. PAPP L. A species new for the Hungarian fauna.

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Rhamphomyia (Pararhamphomyia) tipularia (FALLÉN, 1816) – 1 male: Duna-Ipoly NP: Szokolya, Szén-p. fölött és mellett, 2000. május 13., leg. PAPP L. CHVÁLA and WAGNER (1989) listed it also from Hungary, but we do not know voucher specimen(s) from the modern Hungary. WÉBER (1975) did not include it in his Fauna Hungariae part, not even as a species expected to occur. Chelifera flavella (ZETTERSTEDT, 1838) – 1 male: Zempléni TK: Regéc, Ördög-völgyi patak fölött és mellett, 2000. július 3., leg. PAPP L. WÉBER’s (1975) specimens (1 male, 3 females: Kőszegi-hg., Velemi-erdő, 1960. VII. 11., leg. Mihályi/Zsirkó) were revised and we found that they belong to Ch. trapezina. Consequently this is the first reliable record for this species from Hungary. Dolichocephala engeli NIESIOLOWSKI, 1992 (Dolichocephala engeli: VAILLANT, 1978) – 6 males (leg. L. PAPP): Zempléni TK: Regéc, Ördög-völgy, patak fölött és mellett, 2000. 07. 03.; 1 male: K-Mecsek TK: Komló, Zobákpuszta, Hidasi-völgy, patak fölött, 1999. május 26.; 1 male: Kőszegi TK: Kőszeg, Hármas-p. fölött, mellett, 2000. június 28.; 1 male: A[ggteleki] NP: Szin, Patkós-völgy, 400 m, patakpart, 1988. IX. 12–13.; 2 males: B[ükki] NP: Miskolc, Garadna-völgy, patakpart, 1990. IX. 27. First records for Hungary. Dolichocephala guttata (HALIDAY, 1833) – 9 males (leg. L. PAPP): 1 male: Pécs, Éger-völgy, patak fölött, mellett, 2000. 06. 16.; 1 male: K-Mecsek TK: Komló, Zobákpuszta, Hidasi-völgy, Petasitetum, 1999. május 28.; 4 males: K-Mecsek TK: Óbánya, Óbányai-völgy, patak fölött, 1999. május 25.; 1 male: ibid., patak fölött, mellett, 2000. június 14.; 1 male: [Duna-Ipoly NP] Szendehely, Keskeny-bükki-p. völgye, patakpart [beside the brook], 1997. V. 21.; 1 male: Gerecse TK: Tata-Agostyán, Bocsájtó-völgy [valley], tisztás [clearing], 1990. IX. 7. (plus 10 females of the D. guttata-group in the HNHM). First records for Hungary. Dolichocephala ocellata (COSTA, 1954) – Mecsek Mts, leg. L. PAPP: 1 male: K-Mecsek TK: Komló, Zobákpuszta, Hidasi-völgy, patak fölött, 1999. május 26.; 1 male: ibid., Petasitetum, május 28.; 2 males, 4 females: Melegmányi TT: Pécs, Nagy-mély-v., patak fölött, 1999. május 27.; 1 male: ibid., patak fölött, mellett, 2000. június 15.; 1 male, 1 female: ibid., Melegmányi-völgy, patak fölött, mellett, 2000. június 15.; 1 male, 2 females: K-Mecsek TK: Óbánya, Óbányai-völgy, patak fölött, 1999. május 25/26. First records from Hungary. Dolichocephala thomasi WAGNER, 1984 – 1 male: Zempléni TK [Regéc], Ördög-völgy, Malaise-csapda, 1999. június 8., leg. PAPP L., Szappanos A. This is the first record of this species from Hungary. Clinocera (Kowarzia) barbatula (MIK, 1880) – 1 male: Kőszegi TK: Kőszeg, Hétvezér-forrás [spring], patak fölött, 2000. 07. 25., leg. PAPP L. Another species new for the Hungarian fauna; its closest relative, C. (K.) tenella WAHLBERG, 1844 was published as new to Hungary last year (PAPP & FÖLDVÁRI 2000). Wiedemannia sp. – 1 female: Kács, vízimalom zúgója [at a water-mill], 1955. VIII. 18., leg. Z. Sebess. Its head and most of the legs are lost. This is obviously a Wiedemannia species but it cannot be identified in its damaged state and because of its sex. Otherwise it would be the first reliable record of the Wiedemannia as genus from Hungary. WÉBER (1975) keyed 18 species of this genus from Hungary. However, all but one of their names were published in square brackets (i.e. as species expected to occur). The occurrence of “Eucelidia Zetterstedti Wlk.“ at Budapest, which was published by THALHAMMER (1900) and quoted by Wéber, is improbable, as we (PAPP & FÖLDVÁRI 2000) noted last year. * Acknowledgements – Our sincere thanks are due to Drs MIROSLAV BARTÁK (University of Agriculture, Prague) and RÜDIGER WAGNER (Limnological Stream Station, Schlitz) for their aid in the

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identification of the Rhamphomyia and of the Dolichocephala species, respectively. We also gratefully acknowledge Dr ANDREAS STARK (Ampyx, Halle) for his advice. We especially thank Professor MILAN CHVÁLA (Charles University, Prague) for information he collected for us on the type of Leptopeza rugosiventris STROBL in the STROBL’s collection in Admont, as well as for his advice. This paper was written for the project of “Large blank spots in the Diptera fauna of Hungary”, supported by the Hungarian Scientific Research Fund (OTKA, No. T 30242).

REFERENCES CHILLCOTT, J. G. & TESKEY, H. J. (1983) A revision of the New World genera allied to Megagrapha Melander (Diptera: Empididae). Canad. Ent. 115: 1291–1328. CHVÁLA, M. (1975) The Tachydromiinae (Dipt. Empididae) of Fennoscandia and Denmark. Fauna ent. scand. 3: 1–336. CHVÁLA, M. (1983) The Empidoidea (Diptera) of Fennoscandia and Denmark. II. General part. The families Hybotidae, Atelestidae and Microphoridae. Fauna ent. scand. 12: 1–279. CHVÁLA, M. & WAGNER, R. (1989) Family Empididae. Pp. 228–336. In SOÓS, Á. & PAPP, L. (eds) Catalogue of Palaearctic Diptera, vol. 6. Akadémiai Kiadó, Budapest. COLLIN, J. E. (1961) Empididae. Pp. 1–782. In British Flies, vol. 6. Cambridge Univ. Press, Cambridge. PAPP, L. & FÖLDVÁRI, M. (2000) Empidoidea (Diptera): genera and species new to Hungary. Folia ent. hung. 61: 239–244. SINCLAIR, B. J. & CUMMING, J. M. (2000) Revision of the genus Apterodromia (Diptera: Empidoidea), with a redefinition of the tribe Ocydromiini. Records of the Australian Museum 52: 161–186. STROBL, G. (1910) Die Dipteren von Steiermark. II. Nachtrag. Mitt. naturw. Ver. Steierm. 46(1909): 45–293. THALHAMMER, J. (1900) Diptera, pp. 1–76. In A Magyar Birodalom Állatvilága, Fauna Regni Hungariae vol. 3. A K. M. Természettudományi Társulat, Budapest. [in Latin and Hungarian] WAGNER, R. (1984) Aquatische Empididen (Diptera) aus hessischen Mittelgebirgen und angrenzenden Gebieten. Beitr. Naturkde Dresden 19(1983): 135–146. WÉBER, M. (1975) 28. család: Táncoslegyek–Empididae. Magyarország Állatvilága (Fauna Hungariae) 14(13): 1–220. Akadémiai Kiadó, Budapest. [in Hungarian] WÉBER, M. (1976) Magyarország Empididae (Diptera) fajainak faunakatalógusa. Pécsi Tanárk. Főisk. Tud. Közlem., Ser. 7, Biol. 20: 3–33. [in Hungarian with German summary] WÉBER, M. (1983) Újabb adatok a barcsi borókás Empididae (Diptera) faunájáról [Recent data on the Empididae fauna (Diptera) of the Juniper woodland of Barcs, Hungary]. Dunántúli Dolg. Term. Tud. Sor. Pécs 3: 76–81. WÉBER, M. (1987) The empidid fauna of the Kiskunság National Park (Diptera), pp. 251–257. In MAHUNKA, S. (ed.) The Fauna of the Kiskunság National Park, vol. 2. Akadémiai Kiadó, Budapest. Revised version received 10th September, 2001, accepted 20th December, 2001, published 10th March, 2002

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